adult: August
Last update of
this page: 25-07-05 18:10
adults in the colony
In the Maasvlakte colony many adults try to raise two juveniles. As
soon as the fledglings can move, the family tries to find a more
comfortable place outside the colony, often at the beach or the near fresh
water lakes. By the end of August, 99% of the
adult LBBG from the Maasvlakte and
their juveniles (approximately 70.000 birds) have left the
Maasvlakte and start migrating south (at least leave the immediate feeding
grounds around the colony). For a while, they gather at the beach, in mixed groups of different age-classes.
Several 1.000s of LBBG can be
present up to mid September. In these groups the races graellsii
and intermedius are both present, but hard evidence to categorize
adults to certain races is very difficult, and classification is often only possible when birds
are banded / ringed. From mid-October numbers of LBBG decrease rapidly.
moult in the
primaries
August is normally a month when primary moult is easy to detect, even
on adult birds with folded wings. A small survey of adults in the
Maasvlakte colony did not point to a significant primary moult delay in
adults with begging youngs in the colony. Individual variation in adult LBBG within a
certain colony seems to be at least as large as primary moult variation
between colonies of Dutch intergrades and presumed intermedius from
Scandinavia. In the table below, remaining old primaries are plotted for
mixed groups graellsii, Dutch intergrades and intermedius
at the Maasvlakte in the second half of August. Probably, most birds
originate from the nearby colony and are Dutch intergrades, but large
groups of Scandinavian birds arrive from mid-August onwards.
Intermedius LBBG from Denmark and
southern Norway (intermedius) probably arrive later on the breeding
grounds in spring, compared to Dutch intergrades. Subsequently, it may be
expected that egg-laying date is later in these populations. It remains to
be seen whether or not they show delay in primary moult stage. Such
surveys should only involve ringed (certain origin) adults. At this
moment, such surveys are impossible; comparable analyses are unreliable by
lack of sufficient ring observations.
So far, the hypothesis that intermedius have delay in primary moult
could not be verified, but if it's true, the average remaining old primary
figure in the table is overestimated (if read as applying to Dutch
intergrades) by the input of intermedius arriving in August and
later on. On the other hand, it is expected that Dutch intergrades
with low moult score (arrested or suspended moult) are keen to leave the
Netherlands as soon as possible, leaving a group of adults with relatively
high moult score. At this moment, the figures in the table can be used as
indication for "what average moult stage can be expected in the
second half of August in LBBGs of various origins, present in the
Netherlands".
tables
Remaining
old primaries in adult LBBGs, August 2001. |
|
10 |
9 |
8 |
7 |
6 |
5 |
4 |
3 |
2 |
n: |
m: |
V: |
SD: |
SE: |
15/08: |
6 |
7 |
19 |
21 |
22 |
39 |
18 |
1 |
- |
133 |
6,20 |
2,81 |
1,68 |
0,15 |
21/08: |
13 |
10 |
22 |
32 |
43 |
102 |
49 |
2 |
- |
273 |
5,82 |
2,67 |
1,63 |
0,10 |
28/08: |
14 |
14 |
25 |
28 |
93 |
158 |
139 |
47 |
1 |
519 |
5,22 |
2,54 |
1,59 |
0,07 |
First cell
(10, 15/08 = 6) should be read: on August 15 2001, 6 adults had all 10
primaries old, hence showed no primary moult yet (this is about 4%
on 133 observed adults that day).
n: observations.
m: average.
V (= s2): variance.
SD (= s): standard deviation.
SE (s / sqr n): standard error.
|
Survey
August 16 2001 Maasvlakte, the
Netherlands.
Adult primary score in the colony; adults with begging youngs. |
p3 old |
p4 old |
p5 old |
p6 old |
p7 old |
p8 old |
n: |
average: |
1 |
3 |
3 |
2 |
1 |
1 |
11 |
5,82 |
Survey
of old primaries in adult LBBG on August 22 2002 at Maasvlakte, the
Netherlands (51.57N,04.03E). |
|
Dintelhaven |
Missouriweg |
Papegaaienbek |
p1 old |
2 |
- |
1 |
p2 old |
4 |
1 |
5 |
p3 old |
8 |
- |
8 |
p4 old |
32 |
1 |
10 |
p5 old |
44 |
7 |
14 |
p6 old |
96 |
26 |
12 |
p7 old |
65 |
12 |
9 |
p8 old |
3 |
- |
- |
p9 old |
1 |
- |
- |
p10 old |
- |
- |
- |
old
prims n
old prim m
old prims SD
old prims SE |
255
5.7
1.28
0.08 |
47
6.0
0.92
0.13 |
59
4.7
1.59
0.21 |
Notes:
- For exact location, view the map of Maasvlakte.
- n = observations; m = average;
SD = standard deviation;
SE (s / sqr n) = standard error.
- In the Dintelhaven sample, another 7 birds were scored with
"at least P5 old" in the folded wing, hence P5 or inner
primaries still present. In the extreme case these birds had the
complete primary set old, this would influence the average m
by 0.1.
- old prim m must be read as "primary moult arrived at this
stage for remaining old primaries in average adult LBBG".
- Regarding Dintelhaven: Adult primary score in the colony. Scored adults
at a small pool which is used to bath and preen by adults from the
Dintelhaven colony.
- Regarding Missouriweg: Adult birds feeding begging fledglings in
the colony.
- Regarding Papegaaienbek: A mixed flock of adults preening at the
sandy shores before returning to the Papegaaienbek colony together
with Scandinavian adults using the beaches as stop-over resting
point. |
|
adult LBBG: Dutch intergrade |
LBBG
9cy E246, August
24 2003, Texel, the
Netherlands. P1 missing. |
LBBG 7cy E914, August 18 2001, Missouriweg
Maasvlakte, the
Netherlands. P6 shed, P7-P10 are still old. |
LBBG
9cy EM21, August
15 2004, Missouriweg Maasvlakte, the
Netherlands. Ringed as pullus on July 10 1996. |
LBBG
11cy KR7 August
12 2004, Westkapelle, the Netherlands. Dark individual. Ringed July
07 1993. |
LBBG
9cy EV24
August 12 2004, Westkapelle, the Netherlands. dark individual. Ringed
July 12 1996. |
LBBG
11cy ME9
August 12 2004, Westkapelle, the Netherlands. dark individual. Ringed
July 01 1993. |
LBBG
hybrid =3, August
22 2003, IJmuiden, the
Netherlands. Ringed in the Netherlands in 1999. |
LBBG
6cy AFAF , August
15 2003, Westkapelle, the
Netherlands. |
LBBG
7cy BNAH, August
01 2003, Zeebrugge, Belgium. |
LBBG
7+cy BUAM, August
01 2003, Zeebrugge, Belgium. |
LBBG
adult NLA 5.333.53x, August
28 2003, IJmuiden, the
Netherlands. Ringed in the Netherlands. P8-P10 are still old, P5 slightly
shorter than P4. |
adult LBBG: intermedius |
LBBG
adult SVS 8081001, August
28 2003, IJmuiden, the
Netherlands. Ringed in Sweden. P5-P10 are still old, P2 is fully grown. |
LBBG
adult DKC 4271507 August
12 2004, Westkapelle, the Netherlands. Ringed in Denmark. |
LBBG
adult NOS 4209478 August
12 2004, Westkapelle, the Netherlands. Ringed in Norway. |
LBBG
adult NOS 4111326, August
29 2003, IJmuiden, the
Netherlands. Ringed in Norway. P5-P10 are still old, P3 fully grown. |
LBBG
adult NOS 4163669, August
29 2003, IJmuiden, the
Netherlands. P7-P10 are still old, P3 fully grown. |
LBBG
adult NOS 4191125, August
28 2003, IJmuiden, the Netherlands. P6-P10 are still old, P3 fully grown. |
LBBG
adult NOS 492081, August
11 2003, Boulogne/Mer, France. P5-P10 are still old. All rectrices old. |
adult LBBG: unringed individuals |
Photo
1: LBBG adult & 1cy, August 16 2001, Missouriweg Maasvlakte, the
Netherlands. |
Photo
2: LBBG adult, August 16 2001, Missouriweg Maasvlakte, the
Netherlands. Probably a female with at least p5 to p10 old. |
Photo
3: LBBG adult, August 16 2001, Vuurtorenvlakte Maasvlakte, the
Netherlands. Still no primary moulted: p1-p10 old. |
Photo
4: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the
Netherlands. Adult from the colony; p5 missing, p6 old. |
Photo
5: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the
Netherlands. From the colony, at least the visible primaries are old:
p5-p10. |
Photo
6: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the
Netherlands. Adult from the colony; p5 missing, p6 old. |
Photo
8: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the
Netherlands. P7-p10 old, p6 shed. |
Photo
9: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the
Netherlands. At least p5-p10
are old primaries. |
Photo
10: LBBG adult, August 24 2003,
Texel, the
Netherlands. P1-10 old. |
Photo
11: LBBG adult, August 24 2003, Texel, the
Netherlands. P1-10 old. |
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