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GENETIC DIFFERENTIATION, GEOGRAPHIC VARIATION AND HYBRIDIZATION IN GULLS OF THE LARUS GLAUCESCENS-OCCIDENTALIS COMPLEX
DOUGLAS A. BELL IN: The Condor 98:527-546, 1996
TABLE 1. Locations of the 33 colony areas where specimens of the L. glaucescens-occidentalis complex were collected. Except for L. o. wymani, sample sizes (n) refer to specimens identified a posteriori. Alphanumeric codes correspond to numbered localities in Figure 1. |
Alphanumeric
codes |
Colony area |
L. glaucescens
n |
Hybrids
n |
L.o. occidentalis
n |
L.o. wymani
n |
| BULD1 |
Buldir I., Aleutian Is., AK |
19 |
. |
. |
. |
| KACH2 |
Kachemak Bay, AK |
8 |
. |
. |
. |
| MIDD3 |
Middleton I., AK |
20 |
. |
. |
. |
| MASS4 |
Masset, Queen Charlotte Is., BC |
13 |
2 |
. |
. |
| CUMS5 |
Cumshewa, Queen Charlotte Is., BC |
29 |
4 |
. |
. |
| STJA6 |
St. James I., Queen Charlotte Is., BC |
14 |
1 |
. |
. |
| NVAN7 |
Northern Vancouver I., BC |
9 |
5 |
. |
. |
| GEOR8 |
Georgia Strait, BC |
15 |
6 |
. |
. |
| CLEL9 |
Cleland I., BC |
14 |
13 |
. |
. |
| SJI10 |
San Juan Is., WA |
23 |
17 |
. |
. |
| TAT11 |
Tatoosh I., WA |
14 |
26 |
1 |
. |
| PRO12 |
Protection I., WA |
8 |
8 |
1 |
. |
| DES13 |
Destruction I., WA |
. |
13 |
5 |
. |
| GRA14 |
Grays Harbor, WA |
6 |
24 |
16 |
. |
| PUG15 |
Puget Sound, WA |
27 |
17 |
. |
. |
| WIL16 |
Willapa Bay, WA |
2 |
11 |
4 |
. |
| ESI17 |
East Sand Island, OR |
3 |
21 |
3 |
. |
| TIL18 |
Tillamook, OR |
. |
2 |
12 |
. |
| YAQ19 |
Yaquina Bay, OR |
2 |
8 |
8 |
. |
| COO20 |
Coos Bay, OR |
2 |
3 |
25 |
. |
| CCY21 |
Crescent City, CA |
. |
. |
16 |
. |
| TRN22 |
Trinidad, CA |
. |
. |
16 |
. |
| SUG23 |
Cape Mendocino, CA |
. |
. |
19 |
. |
| MEN24 |
Mendocino, CA |
. |
. |
16 |
. |
| PTR25 |
Point Reyes, CA |
. |
. |
14 |
. |
| FAR26 |
Farallon Is., CA |
. |
. |
27 |
. |
| SFB27 |
San Francisco Bay, CA |
. |
. |
23 |
. |
| ANO28 |
AIio Nuevo I., CA |
. |
. |
6 |
. |
| MOS29 |
Moss Landing, CA |
. |
. |
. |
11 |
| MOR30 |
Morro Bay, CA |
. |
. |
. |
13 |
| ANA31 |
Anacapa I., CA |
. |
. |
. |
32 |
| SNC32 |
San Nicolas I., CA |
. |
. |
. |
19 |
| LCO33 |
Los Coronados I., MX |
. |
. |
. |
10 |
| |
TOTAL: |
228 |
181 |
212 |
85 |
FIGURE 1.
(inset:) Map of 33 colony areas sampled in this study. Numbers correspond to colony alphanumeric codes in Table 1.
(overall:) Map of the hybrid and introgressive zones in the L. glaucescens-occidentalis complex. Pie diagrams illustrate the relative proportions of each morphotype collected at the given colony area. Morphotypes classified a posteriori by DFA. Numbers next to each pie diagram correspond to colony alphanumeric codes (Table 1). The hybrid zone is defined here as the region of breeding sympatry between pure L. glaucescens and L. o. occidentalis, and encompasses colonies 12 (PRO12) to 20 (COO20), excluding 15 (PUG15). Its midpoint is centered around colony 14 (GRA14). The zone of introgression is the region where hybrids were collected as breeding birds, and encompasses colonies 4 (MASS4) to 20.
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Glaucous-winged Gull glaucescens