adult: August

 

Last update of this page: 25-07-05 18:10

adults in the colony

In the Maasvlakte colony many adults try to raise two juveniles. As soon as the fledglings can move, the family tries to find a more comfortable place outside the colony, often at the beach or the near fresh water lakes. By the end of August, 99% of the adult LBBG from the Maasvlakte and their juveniles (approximately 70.000 birds) have left the Maasvlakte and start migrating south (at least leave the immediate feeding grounds around the colony). For a while, they gather at the beach, in mixed groups of different age-classes. 

Several 1.000s of  LBBG can be present up to mid September. In these groups the races graellsii and  intermedius are both present, but hard evidence to categorize adults to certain races is very difficult, and classification is often only possible when birds are banded / ringed. From mid-October numbers of LBBG decrease rapidly. 

moult in the primaries

August is normally a month when primary moult is easy to detect, even on adult birds with folded wings. A small survey of adults in the Maasvlakte colony did not point to a significant primary moult delay in adults with begging youngs in the colony. Individual variation in adult LBBG within a certain colony seems to be at least as large as primary moult variation between colonies of Dutch intergrades and presumed intermedius from Scandinavia. In the table below, remaining old primaries are plotted for mixed groups graellsii, Dutch intergrades and intermedius at the Maasvlakte in the second half of August. Probably, most birds originate from the nearby colony and are Dutch intergrades, but large groups of Scandinavian birds arrive from mid-August onwards. 

Intermedius LBBG from Denmark and southern Norway (intermedius) probably arrive later on the breeding grounds in spring, compared to Dutch intergrades. Subsequently, it may be expected that egg-laying date is later in these populations. It remains to be seen whether or not they show delay in primary moult stage. Such surveys should only involve ringed (certain origin) adults. At this moment, such surveys are impossible; comparable analyses are unreliable by lack of sufficient ring observations.
So far, the hypothesis that intermedius have delay in primary moult could not be verified, but if it's true, the average remaining old primary figure in the table is overestimated (if read as applying to Dutch intergrades) by the input of intermedius arriving in August and later on. On the other hand, it is expected that Dutch intergrades with low moult score (arrested or suspended moult) are keen to leave the Netherlands as soon as possible, leaving a group of adults with relatively high moult score. At this moment, the figures in the table can be used as indication for "what average moult stage can be expected in the second half of August in LBBGs of various origins, present in the Netherlands".

tables

Remaining old primaries in adult LBBGs, August 2001.
  10 9 8 7 6 5 4 3 2 n: m: V: SD: SE:
15/08: 6 7 19 21 22 39 18 1 - 133 6,20 2,81 1,68 0,15
21/08: 13 10 22 32 43 102 49 2 - 273 5,82 2,67 1,63 0,10
28/08: 14 14 25 28 93 158 139 47 1 519 5,22 2,54 1,59 0,07
First cell (10, 15/08 = 6) should be read: on August 15 2001, 6 adults had all 10 primaries old, hence showed no primary moult yet (this is about 4% on 133 observed adults that day).
n: observations.
m: average.
V (= s2): variance.
SD (= s): standard deviation.
SE (s / sqr n): standard error.

 

Survey August 16 2001 Maasvlakte, the Netherlands. Adult primary score in the colony; adults with begging youngs.
p3 old p4 old p5 old p6 old p7 old p8 old n: average:
1 3 3 2 1 1 11 5,82

 

Survey of old primaries in adult LBBG on August 22 2002 at Maasvlakte, the Netherlands (51.57N,04.03E).
  Dintelhaven Missouriweg Papegaaienbek
p1 old 2 - 1
p2 old 4 1 5
p3 old 8 - 8
p4 old 32 1 10
p5 old 44 7 14
p6 old 96 26 12
p7 old 65 12 9
p8 old 3 - -
p9 old 1 - -
p10 old - - -
old prims n
old prim m
old prims SD
old prims SE
255
5.7
1.28
0.08
47
6.0
0.92
0.13
59
4.7
1.59
0.21

Notes:
- For exact location, view the map of Maasvlakte.
- n = observations; m = average; SD = standard deviation; SE (s / sqr n) = standard error.
- In the Dintelhaven sample, another 7 birds were scored with "at least P5 old" in the folded wing, hence P5 or inner primaries still present. In the extreme case these birds had the complete primary set old, this would influence the average m by 0.1.
- old prim m must be read as "primary moult arrived at this stage for remaining old primaries in average adult LBBG".
- Regarding Dintelhaven: Adult primary score in the colony. Scored adults at a small pool which is used to bath and preen by adults from the Dintelhaven colony.
- Regarding Missouriweg: Adult birds feeding begging fledglings in the colony.
- Regarding Papegaaienbek: A mixed flock of adults preening at the sandy shores before returning to the Papegaaienbek colony together with Scandinavian adults using the beaches as stop-over resting point.

 


adult LBBG: Dutch intergrade
ad lbbg in August, ringed in the Netherlands. (75385 bytes)LBBG 9cy E246, August 24 2003, Texel, the Netherlands. P1 missing.
ad LBBG in August, ringed in the Netherlands. (64808 bytes)LBBG 7cy E914, August 18 2001, Missouriweg Maasvlakte, the Netherlands. P6 shed, P7-P10 are still old. 
ad LBBG in August, ringed in the Netherlands. (59747 bytes)LBBG 9cy EM21, August 15 2004, Missouriweg Maasvlakte, the Netherlands. Ringed as pullus on July 10 1996.
ad LBBG in August, ringed in the Netherlands. (84068 bytes)LBBG 11cy KR7 August 12 2004, Westkapelle, the Netherlands. Dark individual. Ringed July 07 1993.
ad LBBG in August, ringed in the Netherlands. (84266 bytes)LBBG 9cy EV24 August 12 2004, Westkapelle, the Netherlands. dark individual. Ringed July 12 1996.
ad LBBG in August, ringed in the Netherlands. (72478 bytes)LBBG 11cy ME9 August 12 2004, Westkapelle, the Netherlands. dark individual. Ringed July 01 1993.
ad LBBG in August, ringed in the Netherlands. (87390 bytes)LBBG hybrid =3, August 22 2003, IJmuiden, the Netherlands. Ringed in the Netherlands in 1999.
sub-adult LBBG in August, ringed in Belgium. (53906 bytes)LBBG 6cy AFAF , August 15 2003, Westkapelle, the Netherlands. 
sub-adult LBBG in August, ringed in Belgium. (76270 bytes)LBBG 7cy BNAH, August 01 2003, Zeebrugge, Belgium.
sub-adult LBBG in August, ringed in Belgium. (57738 bytes)LBBG 7+cy BUAM, August 01 2003, Zeebrugge, Belgium.
ad LBBG in August, ringed in the Netherlands. (57118 bytes)LBBG adult NLA 5.333.53x, August 28 2003, IJmuiden, the Netherlands. Ringed in the Netherlands. P8-P10 are still old, P5 slightly shorter than P4.

adult LBBG: intermedius
ad LBBG in August, ringed in Sweden. (56368 bytes)LBBG adult SVS 8081001, August 28 2003, IJmuiden, the Netherlands. Ringed in Sweden. P5-P10 are still old, P2 is fully grown.
ad LBBG in August, ringed in Denmark. (83984 bytes)LBBG adult DKC 4271507 August 12 2004, Westkapelle, the Netherlands. Ringed in Denmark.
ad LBBG in August, ringed in Norway. (66598 bytes)LBBG adult NOS 4209478 August 12 2004, Westkapelle, the Netherlands. Ringed in Norway.
ad LBBG in August, ringed in Norway. (87905 bytes)LBBG adult NOS 4111326, August 29 2003, IJmuiden, the Netherlands. Ringed in Norway. P5-P10 are still old, P3 fully grown.
ad LBBG in August, ringed in Norway. (73700 bytes)LBBG adult NOS 4163669, August 29 2003, IJmuiden, the Netherlands. P7-P10 are still old, P3 fully grown.
ad LBBG in August, ringed in Norway. (66816 bytes)LBBG adult NOS 4191125, August 28 2003, IJmuiden, the Netherlands. P6-P10 are still old, P3 fully grown.
ad LBBG in August, ringed in Norway. (64193 bytes)LBBG adult NOS 492081, August 11 2003, Boulogne/Mer, France. P5-P10 are still old. All rectrices old.

adult LBBG: unringed individuals
Photo 1: LBBG adult & 1cy, August 16 2001, Missouriweg Maasvlakte, the Netherlands.
5789.jpg (87878 bytes)Photo 2: LBBG adult, August 16 2001, Missouriweg Maasvlakte, the Netherlands. Probably a female with at least p5 to p10 old.
5810.jpg (72864 bytes)Photo 3: LBBG adult, August 16 2001, Vuurtorenvlakte Maasvlakte, the Netherlands. Still no primary moulted: p1-p10 old.
5880.jpg (77357 bytes)Photo 4: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the Netherlands. Adult from the colony; p5 missing, p6 old.
5882.jpg (82268 bytes)Photo 5: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the Netherlands. From the colony, at least the visible primaries are old: p5-p10.
5883.jpg (95425 bytes)Photo 6: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the Netherlands. Adult from the colony; p5 missing, p6 old.
5889.jpg (98610 bytes)Photo 8: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the Netherlands. P7-p10 old, p6 shed.
5894.jpg (91853 bytes)Photo 9: LBBG adult, August 18 2001, Missouriweg Maasvlakte, the Netherlands. At least p5-p10 are old primaries. 
ad lbbg in August. (66207 bytes)Photo 10: LBBG adult, August 24 2003, Texel, the Netherlands. P1-10 old.
ad lbbg in August. (64438 bytes)Photo 11: LBBG adult, August 24 2003, Texel, the Netherlands. P1-10 old.