third
calendar year: August
Last update of
this page: 27-10-03 02:20
In
LBBG nearly all feathers are being replaced now. There is a great
difference between the start and the end of August. Birds change into a
fresh bird with adult-like upper parts and under parts. Wing coverts and
tertials become 3rd generation and mantle and scapulars become 4th
or 5th generation, after the moult to 3rd winter plumage. From head to
toe you can see the following changes.
- The
bill becomes paler. Variation in bill colour is low. Bills with a
black base (present in 9%, n= 100) are rare while some advanced
birds (17%) show an adult-like bill. On average, birds show
an adult bill with a dark smudge on the upper mandible (74%).
- The
iris turns pale yellow, but a white or darker peppered iris occur.
- The head varies from all-white without any speckling (old, worn,
15%), finely streaked from crown to neck (65%) to new heavily streaked
(20%). Birds actively moulting the head-feathers look spiky.
- The under-parts are
actively moulted into white feathers. Nearly 100% of
the birds shows all-white under-parts, a few have streaking
in the hind-neck.
- By the end of August,
all 3cy LBBGs show adult grey
upper-parts.
- Many to nearly all 3cy LBBGs
have finished moult of
mantle and scapulars (by September all birds). The upper-parts
resemble adults now (being 4th or 5th generation).
- Tertials on 3cy LBBGs in August can exist of
old
2nd generation (at least) feathers or newly moulted 3rd generation feathers. The
difference is obvious: the new tertials resemble adults,
contrary to the blackish centred old tertials. About 50% of 3cy LBBGs
have all
tertials new, with new upper tertials and old lower tertials being second
best.
- The wing-coverts are in active moult, so a gap in the
covert bars is very
common.
The moult starts with the medians.
When median covert moult is halfway, birds start moulting
the greater and lesser coverts as well, normally starting with inner and outer greaters. Last moulted feathers are the outer
lessers.
- Common
moult-state in 3cy LBBGs by late August is:
new inner and central medians (3rd generation), 4 new inner greaters (3rd
generation), missing the
central greaters and growing the outer-half inwards and 50% of the
lessers old. The 3rd generation coverts normally are adult-grey without barring.
- Primaries are moulted outwards, the primary-score
by the end of
August averages 34 (n = 26), all birds showing fully grown p4 at least
(see table). Third generation primaries are very adult-like but p10 often shows an obvious
smaller mirror. (Later in the year, bleaching gives a browner tinge on the primaries
than on adults.)
- Secondaries are dropped inwards, from the moment
p5 or p6 is halfway grown, creating a gap from p1 inwards. Only a very few 3cy show secondary-moult in August. No or nearly none 3cy
show fully 3rd generation secondaries by the end of August.
- The
under-wing becomes paler, under-wing coverts are replaced by
adult-like feathers. The under-wing coverts are probably moulted in
the same sequence as the upper-wing coverts.
-
The
tail is old, 2nd generation on many birds (60%), in active moult (20%) and
fully grown 3rd generation, white or with very restricted black
markings (20%). Old worn tail-feathers occur as some 3cy and 4cy LBBGs may moult the tail on the wintering
grounds, subsequently returning with a new tail in spring.
-
Leg
colour varies from pink to (pale) yellow.
-
Birds
with advanced mantle and scapulars already look exactly like 4cy birds
as they return by May in the Netherlands.
Tables
|
remaining
old primaries |
full-grown
new primaries |
|
#
of new tertials |
|
0 |
1 |
2 |
3 |
4 |
5 |
6 |
p2 |
p3 |
p4 |
p5 |
p6 |
p7 |
|
0 |
1 |
2 |
3 |
4 |
5 |
6 |
15/8 |
|
|
2 |
10 |
10 |
3 |
2 |
2 |
4 |
9 |
10 |
3 |
|
|
|
|
|
|
|
|
1 |
21/8 |
|
|
2 |
7 |
6 |
3 |
|
|
4 |
6 |
6 |
2 |
|
|
1 |
|
2 |
5 |
1 |
2 |
4 |
23/8 |
1 |
2 |
12 |
8 |
5 |
|
|
|
1 |
4 |
13 |
9 |
1 |
|
|
|
3 |
5 |
5 |
3 |
12 |
28/8 |
2 |
4 |
11 |
6 |
3 |
|
|
|
|
5 |
7 |
8 |
6 |
|
|
1 |
1 |
3 |
10 |
|
11 |
31/8 |
2 |
7 |
14 |
9 |
4 |
|
|
|
|
6 |
14 |
11 |
5 |
|
|
|
3 |
5 |
12 |
5 |
11 |
|
0: no old primaries
left,
1: only p10 is still old,
6: p5 still 2nd generation. |
p2:
only p1-p2 new, others missing or old,
p7:
p1-p7 replaced 3rd generation. |
|
|
lesser
coverts |
|
median
coverts |
|
greater
coverts |
|
0 |
1 |
2 |
3 |
4 |
5 |
0 |
1 |
2 |
3 |
4 |
5 |
0 |
1 |
2 |
3 |
4 |
5 |
15/8 |
1 |
|
|
|
|
|
|
|
|
|
|
1 |
|
1 |
|
|
|
|
21/8 |
4 |
3 |
4 |
4 |
1 |
2 |
|
|
|
2 |
1 |
15 |
1 |
4 |
9 |
1 |
1 |
2 |
23/8 |
7 |
4 |
3 |
4 |
4 |
6 |
|
|
|
|
4 |
24 |
|
7 |
6 |
2 |
8 |
5 |
28/8 |
2 |
1 |
2 |
5 |
9 |
7 |
|
|
|
|
4 |
22 |
|
4 |
3 |
2 |
9 |
8 |
31/8 |
|
4 |
10 |
5 |
8 |
9 |
|
|
|
|
5 |
31 |
|
5 |
5 |
4 |
9 |
13 |
Coverts
score with
0 = all old, 1 = 25% new, 2 = 50% new, 3 = 75% new,
4 = 99%
new, 5 = 100% new. |
Primary
moult score of 3cy LBBG, at Naaktstrand, Maasvlakte, the Netherlands
(51.59N,
04.02E) on August 14 2003. |
. |
P3
fg |
P4
fg |
P5
fg |
P6
fg |
P8
fg |
n: |
P6-P10 |
2 |
- |
- |
- |
- |
2 |
P7-P10 |
- |
7 |
- |
- |
- |
7 |
P8-P10 |
- |
1 |
10 |
1 |
- |
12 |
P9-P10 |
- |
- |
- |
14 |
1* |
15 |
n: |
1 |
8 |
10 |
15 |
1 |
36 |
*
arrested moult at P8. P1 fg 4th gen, P2 growing, P3 missing, P4-P8
3rd gen, P9-P10 2nd gen.
n = 35; m old P = 1.7; SD old P = 0.95.
P8-P10 = P8-P10 are still old and present.
Further notes: see previous table. |
Primary
moult score in 3cy LBBG, at Westkapelle, the Netherlands (51.33N,
03.25E) on August 15 2003. |
. |
P3
fg |
P4
fg |
P5
fg |
P6
fg |
P7
fg |
n: |
P6-P10 |
1 |
- |
- |
- |
- |
1 |
P7-P10 |
- |
1 |
1 |
1 |
- |
3 |
P8-P10 |
- |
- |
5 |
- |
- |
5 |
P9-P10 |
- |
- |
- |
5 |
- |
5 |
P10
old |
- |
- |
- |
- |
1 |
1 |
n: |
1 |
1 |
6 |
6 |
1 |
15 |
For
notes: see previous table.
Survey group of 82 LBBG, including 15 3cy. |
primary
moult score in 3cy LBBG, at Maasvlakte, the Netherlands, (51.59N,
04.02E), August
22 2002. |
. |
P6-P10 |
P7-P10 |
P8-P10 |
P9-P10 |
P10 old |
no
old P |
n: |
p7
fg |
- |
- |
- |
1 |
3 |
3 |
7 |
p6
fg |
- |
- |
5 |
7 |
- |
- |
12 |
p5
fg |
- |
3 |
12 |
- |
- |
- |
15 |
p4
fg |
2 |
1 |
- |
- |
- |
- |
3 |
?
fg |
- |
2 |
1 |
- |
- |
- |
3 |
n: |
2 |
6 |
18 |
8 |
3 |
3 |
40 |
n =
40; m old P = 2.7; SD old P = 1.21.
P6-P10 = P6-P10 are still old and present.
? fg = fully grown not known. Fully grown primary score most
probably P6 or less.
fg = fully grown. In our field surveys, a primary is 'fully grown'
when it exceeds the previous feather in the folded wing. Actually,
such 'fully grown' feathers may have to grow a little, but under
field circumstances, it's very difficult to find out if the waxy
sheath, which is the standard measurement for fully grown primaries,
is still present. |
primary
moult score in 3cy LBBG, IJmuiden, the Netherlands (52.27N, 04.33E), August
30 2002. |
. |
P7-P10 |
P8-P10 |
P9-P10 |
P10 old |
no
old P |
n: |
p8
fg |
- |
- |
1* |
- |
- |
1 |
p7
fg |
- |
- |
2 |
6 |
6 |
14 |
p6
fg |
- |
4 |
9 |
- |
- |
13 |
p5
fg |
- |
4 |
- |
- |
- |
4 |
p4
fg |
1 |
- |
- |
- |
- |
1 |
n: |
1 |
8 |
12 |
6 |
6 |
33 |
n =
33; m old P = 1.8; SD old P = 1.12; m new P = 6.3; SD new P = 0.85.
* = arrested moult, P1-P8 moulted earlier on the wintering grounds.
Further notes: see previous table. |
|
LBBG intermedius
J4A3
3cy July 25 & August 21 2003, IJmuiden, the Netherlands.
LBBG Dutch
intergrade L-9833?
3cy, August 01 2003, Zeebrugge, Belgium.
Photo
1: LBBG 3cy August 23 2000, Le Portel, France.
Photo 2: LBBG 3cy, August 18 2001,
Missouriweg Maasvlakte, the Netherlands.
Photo
3: LBBG 3cy, August 18 2001, Missouriweg Maasvlakte, the Netherlands.
Photo
4: LBBG 3cy, August 16 2001, Vuurtorenvlakte Maasvlakte, the
Netherlands.
Photo 5: LBBG 3cy, August 22 2002, Papegaaienbek, Maasvlakte, the
Netherlands.
ARRESTED MOULT IN 3CY
LBBG
LBBG 3cy August
29 2003, IJmuiden, the Netherlands. Probably intermedius, arrested
moult at P8.
|