Lesser Black-backed Gull (graellsii & intermedius)

(last update: 14-11-2017 )

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LBBG rings

lbbg 1cy May
lbbg 1cy June
lbbg 1cy July
lbbg 1cy August
lbbg 1cy September
lbbg 1cy October
lbbg 1cy November
lbbg 1cy December

lbbg 2cy January
lbbg 2cy February
lbbg 2cy March
lbbg 2cy April
lbbg 2cy May
lbbg 2cy June
lbbg 2cy July
lbbg 2cy August
lbbg 2cy September
lbbg 2cy October
lbbg 2cy November
lbbg 2cy December

lbbg 3cy January
lbbg 3cy February
lbbg 3cy March
lbbg 3cy April
lbbg 3cy May
lbbg 3cy June
lbbg 3cy July
lbbg 3cy August
lbbg 3cy September
lbbg 3cy October
lbbg 3cy November
lbbg 3cy December

lbbg sub-ad January
lbbg sub-ad February
lbbg sub-ad March
lbbg sub-ad April
lbbg sub-ad May
lbbg sub-ad June
lbbg sub-ad July
lbbg sub-ad August
lbbg sub-ad Sept
lbbg sub-ad Oct
lbbg sub-ad Nov
lbbg sub-ad Dec

lbbg adult January
lbbg adult February
lbbg adult March
lbbg adult April
lbbg adult May
lbbg adult June
lbbg adult July
lbbg adult August
lbbg adult September
lbbg adult October
lbbg adult November
lbbg adult December

adult: June

 

At the Maasvlakte colonies in the Netherlands (see map), eggs are hatched by the first week of June and many nests contain three nestlings (breeding season 2001 and 2002). Pulli can be found everywhere. First data in 2001 estimate the colonies to be expanded by over 20%, now approximately 40,000 nests at the Europoort location alone (greater Rotterdam district). New colonies arise at plots which are still not occupied by industrial companies. 

In this picture you see female E832, ringed at the nest at Maasvlakte July 1995, now in her 6th summer and raising three pulli. Normally the elder adults occupy the sites used last year(s), as these birds know safety, feeding routes and places, etc. New colonies seem to be occupied by younger adults, as some ring-data suggest. Older LBBGs disturbed from 'ancient' colonies seem to move less far from the places they were forced to abandon. It's interesting to find out whether new colonies abroad are started by relatively young adults.

Moult:

Raising pulli is very time consuming and all moult in the wing-coverts is suspended in adults.
In some individuals, head feather moult is started mid-June and only the odd individuals moult coverts at a low pace. 

Counting back from the results of surveys done in August 2001, a small proportion of adult LBBG must have shed inner primaries by June. Last years, we did not collect sufficient material to estimate primary moult scores for this age class in June, so a small survey on 200 adult birds was done. We scored preening and bathing adults from the near colony (several 1.000's breeding pairs). The first estimation was done on arriving birds, checking the age while bathing and check primary moult score again at departure. The table shows the results: 75% of local (!) adult LBBG have still all the primaries old (p1-p10 old) and a small minority of about 20% has started moult with dropping p1.

Tables & Graph

starting date: 28 May 04 Jun 11 Jun 18 Jun 25 Jun 02 Jul
week no: 22 23 24 25 26 27
REM 0 0 0 0 0 0 0
REM 1 0 0 0 0 0 0
REM 2 0 0 0 0 0 0
REM 3 0 0 0 0 0 0
REM 4 0 0 0 0 0 0
REM 5 0 0 0 0 0 0
REM 6 0 0 0 0 0 1
REM 7 0 0 0 1 0 14
REM 8 5 3 3 13 0 64
REM 9 12 6 12 45 1 112
REM 10 102 17 28 169 5 127
0 0
n = 119 26 43 228 6 318
average P = 9.8 9.5 9.6 9.7 9.8 9.1
95% CID = 0.09 0.29 0.19 0.08 0.43 0.10
inverse = 1.98 2.06 2.02 1.97 2.57 1.97
Variantie (s2) = 0.24 0.50 0.39 0.36 0.17 0.80
SD (s) = 0.49 0.71 0.63 0.60 0.41 0.89
SE (sx) = 0.04 0.14 0.10 0.04 0.17 0.05
% class:
0 0 0 0 0 0 0
1 0 0 0 0 0 0
2 0 0 0 0 0 0
3 0 0 0 0 0 0
4 0 0 0 0 0 0
5 0 0 0 0 0 0
6 0 0 0 0 0 0
7 0 0 0 0 0 4
8 4 12 7 6 0 20
9 10 23 28 20 17 35
10 86 65 65 74 83 40
% control: 100 100 100 100 100 100

Table above, data for the Netherlands (2000-2010), results for June (aggregated week): number of remaining old primaries in adults.
REM10 = 10 primaries still old (hence, complete moult not started yet).

Above: Scatter Plot for PMS in adult Lesser Black-backed Gulls. Number of remaining old primaries throughout the year (arrested moult excluded). Sample size is 6.171 birds.

Above: Scatter Plot for PMS in adult Lesser Black-backed Gulls. Number of remaining old primaries for June (arrested moult excluded). Sample size for this graph is 339 ringed birds + 401 unringed birds.

starting date:

21 May

04 Jun

02 Jul

week no:

21

23

27

NEW 0

92

8

1

NEW P1

4

0

1

NEW P2

0

0

0

NEW P3

0

0

0

NEW P4

0

0

0

NEW P5

0

0

0

NEW P6

0

0

0

NEW P7

0

0

0

NEW P8

0

0

0

NEW P9

0

0

0

NEW P10

0

n =

96

8

2

average NEW P =

0.0

0.0

0.5

95% CID =

0.04

0.00

6.35

inverse =

1.99

2.36

12.71

Variantie (s2) =

0.04

0.00

0.50

SD (s) =

0.20

0.00

0.71

SE (sx) =

0.02

0.00

0.50

% class

0

96

100

50

1

4

0

50

2

0

0

0

3

0

0

0

4

0

0

0

5

0

0

0

6

0

0

0

7

0

0

0

8

0

0

0

9

0

0

0

10

0

0

0

% control:

100

100

100

Table above, data for the Netherlands (2000-2010), results for June (aggregated week): number of new, fully grown primaries in adults.
NEW0 = no new primaries visible (arrested moult excluded in this analysis). So either P1 still old, or P1 dropped, but the very tip still not visible.
NEW1 = P1 longest new primary. Range between "at least the tip of P1 is visible" until "tip of P2 still shorter than P1".
NEW2 = P2 longest new primary. NEW3 = etc.

Above: Scatter Plot for PMS in adult Lesser Black-backed Gulls throughout the year. Longest new primary is classified "fully grown". Sample size is 2.969 birds. Birds scored on NEW primaries (arrested moult excluded).

Above: Scatter Plot for PMS in adult Lesser Black-backed Gulls. Scoring on longest new primary, which is classified "fully grown" for June (arrested moult excluded). Sample size for this graph is 10 ringed birds.

The earliest date for birds with missing primaries is 30 April 2006, which is not the earliest day in our complete sample. We scored 98 Lesser Black-backed Gulls prior to that date, some already in February. As can be expected, none of these was in primary moult, and this group was excluded in our further analysis as it will only result in a misfit for the onset of moult, using linear regression statistics.

The last scoring days were in November, with a larger sample (133 adults) on 04 November 2010. From that date we have the last birds with retaining old primaries (13 birds with 1, 2 or even 3 old primaries; 120 birds with no old primaries left). From other days in November, only 10 birds were scored (all ringed adults). They all had no old primaries left.

The first tine a new primary (P1) was visible, was on 08 May 2006. Prior to this date, birds were scored missing inner primaries, but the very tip was still not visible (and therefore not scored). As soon as the tip of P1 was visible, this was automatically scored "fully grown" For the other primaries to be scored fully grown, they had to exceed the previous primary in length (so P2 fully grown, only from when it is longer than P1).

Peter Stewart presents onset of primary moult by scores for caught adult Lesser Black-backed Gulls in the Severn Estuary, U.K., repeated in the table below. To read his results, see this PDF file.  

Table: the number of old remaining primaries present. Compare PMS of adult graellsii at half monthly intervals. Data from the Severn Estuary Gull Group by Peter Stewart: The primary moult of the Lesser Black-backed Gull (special publication No 1).

 

Netherlands

Severn Estuary U.K.

 

PMS range

N:

Average PMS

PMS range

N:

Average PMS

1-15 apr

10

49

10

8-10

216

9,9

16-30 apr

10

47

10

10

197

10,0

1-15 may

8-10

849

9,8

8-10

167

9,9

16-31 may

7-10

380

9,7

7-10

76

9,5

1-15 jun

8-10

104

9,6

7-10

33

8,6

16-30 jun
7-10
253
9,7
6-10
65
8,2
1-15 jul
6-10
605
9,2
5-10
176
7,4

LINEAR REGRESSION

Onset of primary moult can best be illustrated by a sinusoid for the first few weeks and last weeks, but there appears to be linear dependency over much of the central moult period.

LINEAR REGRESSION FOR OLD REMAINING PRIMARIES

Complete sample: Linear regression (n = 6171, R2= 0.820, sign: P<0,000)
PMS = -0,381* WEEK + 17,967

Ringed adults: Linear regression (n = 2069, R2= 0.800, sign: P<0,000)
PMS = -0,325* WEEK + 16,270

Unringed adult: Linear regression (n = 4102, R2= 0.779, sign: P<0,000)
PMS = -0,513* WEEK + 23,031

LINEAR REGRESSION FOR NEW FULLY GROWN PRIMARIES

Complete sample: Linear regression (n = 2969, R2= 0.930, sign: P<0,000)
PMS = 0,302* WEEK - 5,933

Ringed adults: Linear regression (n = 1209, R2= 0.888, sign: P<0,000)
PMS = 0,252* WEEK - 4,747

Unringed adult: Linear regression (n = 1760, R2= 0.833, sign: P<0,000)
PMS = 0,442* WEEK - 11,906

Primary moult score of adult LBBG June 18 2002, at Dintelhaven colony, Maasvlakte, the Netherlands (51.59N,04.02E).
p3 shed 1
p2 shed 11
p1 shed 43
0 shed 145
n = 200.
two birds had p2 shed, while p1 was (almost) fully grown, hence followed a "condensed" moult strategy.

 

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U.K.: graellsii.
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Lesser Black-backed Gull graellsii RXB 5cy, 6cy, 9cy, 10cy, May 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
adult LBBG in June, ringed in Britain. (61948 bytes)Lesser Black-backed Gull graellsii BTO FA 8370? adult, June 02 2003, Papegaaienbek, Maasvlakte, the Netherlands. An adult ringed with a BTO ring: London FA 8370?.
Lesser Black-backed Gull graellsii BTO FV 38253 8cy-9cy, June 2013 & April 2014, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii BTO GG49760 19cy & 21cy, 2012 & 2014, Calais, NW France. Picture: Jean-Michel Sauvage.
adult LBBG in June, ringed in the U.K. (49422 bytes)Lesser Black-backed Gull graellsii BTO GT 75716 adult, June 02 2003, Papegaaienbek, Maasvlakte, the Netherlands.
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France: graellsii.
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Lesser Black-backed Gull graellsii 255K 6cy, May-June 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii 377N 5cy, June 23 2010, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii 399N 8cy, April, May & June 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii 452N 1,5,7cy, June 24 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii 522K 2CY & 6CY, April 2010 & June 2014, Spain & the Netherlands. Picture: Antonio Gutierrez & Merijn Loeve.
Lesser Black-backed Gull graellsii 726N 6cy-7cy, June 2013 & April 2014, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii 989N 5CY & 8CY, 2010 & 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii FRP EA558292 adult, June 28 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
Ringed as adult.
Lesser Black-backed Gull graellsii l FRP EA639841 / 085N 8CY-9CY, June 25 2012 & April 26 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii FRP EA677791 adult, April, May & June 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii FRP EA67799x adult, June 28 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
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Belgium: graellsii 'Dutch intergrade'.
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juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii HAAB 11CY, May - June 2016, Maasvlakte (Amazonehaven), the Netherlands (51.56 N 04.03 E).
ad LBBG in Januari-April, ringed in the Netherlands. (76807 bytes)Lesser Black-backed Gull graellsii HMAL 7CY, June 03 2014, Nieuwdorp waste dump - Vlissingen, the Netherlands. Picture: Merijn Loeve.
Lesser Black-backed Gull graellsii KJAM 6cy, May, June & July 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii MNAY 8CY, June 13 2016, Katwijk, the Netherlands.
Lesser Black-backed Gull graellsii NTAZ 4cy, April & June 2013, Calais, NW France. Picture: Jean-Michel Sauvage.
adult LBBG in June, ringed in Belgium. (74230 bytes)Lesser Black-backed Gull graellsii BLB L-92594 adult, June 17 2002, EMO - Maasvlakte, the Netherlands.
Belgium ring, but wrong reading of code. In the colonies at the Maasvlakte different birds with British and Belgium rings are found breeding with local Dutch intergrades.
Lesser Black-backed Gull graellsii BLB L-900238 10CY-11CY, June 2013 & April 2014, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii BLB L-902113 8CY-11CY, April 2012 - April 2015, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii BLB L-906179 6cy-10cy, 2010-2014, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii BLB L-906775 6CY-7CY, 9CY, April 2012 & April 2013, June 2015, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii BLB L-912000 6CY, April & June 2014, Calais, NW France. Picture: Jean-Michel Sauvage.
adult LBBG in June, ringed in Belgium. (57581 bytes)Lesser Black-backed Gull graellsii BLB L-?9951, June 02 2003, Dintelhaven, Maasvlakte, the Netherlands.
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the Netherlands: graellsii 'Dutch intergrade'.
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Lesser Black-backed Gull graellsii OV 6cy-10cy, 2010-2014, Calais, NW France. Picture: Jean-Michel Sauvage.
Lesser Black-backed Gull graellsii CM adult, June 01 2015, Calais, NW France. Picture: Jean-Michel Sauvage.
2275.jpg (40492 bytes)Lesser Black-backed Gull graellsii DJ 8CY-9CY & 11CY, 2014-2015 & 2017, Maasvlakte & Utrecht, the Netherlands. Picture: Merijn Loeve, Herman Bouman & Mars Muusse.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii P2 10CY, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii 6.9 6CY, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii T+ adult, June 25 2013, Groningen, the Netherlands. Picture: Henri Zomer.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii F||0 adult, June 18 2016, Katwijk, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii F||V adult, June 18 2016, Katwijk, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii F||W adult, June 18 2016, Katwijk, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii F||Y adult, June 18 2016, Katwijk, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii H||0 adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii H||7 adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii H||8 adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii H||9 adult, June 2016 & May 2018, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii H||Y adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii B8 adult, June 08 2015, Maasvlakte - Europoort, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii BW adult, March 26 & June 08 2015, Maasvlakte - Europoort, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii BX adult, April 10 & June 08 2015, Maasvlakte - Europoort, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii C.V adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii EA (formerly EY98) 18CY, May 23 & June 02 2014, Maasvlakte, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes) Lesser Black-backed Gull graellsii EK adult, June-July 2015, Rotterdam & Tilburg waste dump, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii E.Z adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii 2N 5CY, June - July 2016, Boulogne-sur-Mer, NW France. Picture: Jean-Michel Sauvage.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii 4.7 5CY, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii C3 adult, May-June 2015, Maasvlakte - Europoort, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii C8 adult, June 08 2015, Maasvlakte - Europoort, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii 9.H 5CY, June 06 2016, Katwijk, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii BH adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii BN adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii BP adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii GR adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii HC 5CY, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii KR adult, June 08 27 2015, Maasvlakte - Europoort, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii M6 adult, June 08 27 2015, Maasvlakte - Europoort, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii NW adult, May 2015 & June 2016, Katwijk, the Netherlands. Picture: Merijn Loeve.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii T9 5CY, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii TG 5CY, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii 40 adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii 41 adult, June 06 2016, Maasvlakte, the Netherlands.
ad LBBG in Januari-April, ringed in the Netherlands. (94477 bytes)Lesser Black-backed Gull graellsii 56 adult, June 2016 & May 2018, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii N3 (was E556 E), 16CY, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii NE adult, August 2014 & May-June 2016, Boulogne sur mer, NW France & Maasvlakte, the Netherlands. Picture: Jean-Michel Sauvage & Mars Muusse.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii VW adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii W8 adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii W+ adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii YU adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii ZB adult, June 06 2016, Maasvlakte, the Netherlands.
Lesser Black-backed Gull graellsii P.3 5CY, June-July 2017, Katwijk, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii E= adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii TP (was TH5), 20CY, June 20 2016, Katwijk, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii VL (was EP05), 21CY, June 06 2016, Maasvlakte, the Netherlands.
Lesser Black-backed Gull graellsii E C77 10CY-11CY, April 2012 & June 2013, Maasvlakte & Dordrecht, the Netherlands. Picture: Merijn Loeve & Maarten van Kleinwee.
adult LBBG in June, ringed in the Netherlands. (75836 bytes)Lesser Black-backed Gull graellsii K98 12cy, June 17 2002, Papegaaienbek - Maasvlakte, the Netherlands. Ringed at Maasvlakte as pullus, July 04 1991.
adult LBBG in June, ringed in the Netherlands. (68885 bytes)Lesser Black-backed Gull graellsii T88 12cy, June 17 & 18 2002, Papegaaienbek - Maasvlakte, the Netherlands. Ringed at Maasvlakte as pullus, July 08 1991.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)Lesser Black-backed Gull graellsii TN4 14CY & 16CY, April 2011 & June 2013, the Netherlands. Picture Ruud Altenburg & Maarten van Kleinwee.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii FADU 8+cy , June 24 2011, Texel, the Netherlands. Ringed as breeding female.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii FAHP 18cy, June 24 2011, Texel, the Netherlands. Ringed as breeding female.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii FAHR 6+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding female.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii FAKT 5+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding female.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii KLAS 5cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MADB 6+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MAFZ 15cy, June 24 2011, Texel, the Netherlands. Ringed as breeding female.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MAKL 7+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MAKR 7+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MALZ 5+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MAMP 6+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MAMZ 6+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MAPJ 4+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MAPV 4+cy, June 24 2011, Texel, the Netherlands. Ringed as breeding male.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii MARS adult, June 8 2012, Texel, the Netherlands. Ringed as breeding adult.
adult LBBG in June, ringed in the Netherlands. (57378 bytes)Lesser Black-backed Gull graellsii CY5 10cy, June 17 2002, Papegaaienbek Maasvlakte, the Netherlands.
Ringed June 18 1993. Head feather moult has started around the eye.
adult LBBG in June, ringed in the Netherlands. (62470 bytes)Lesser Black-backed Gull graellsii MA9 10cy, June 17 2002, Papegaaienbek Maasvlakte, the Netherlands.
Ringed July 01 1993.
adult LBBG in June, ringed in the Netherlands. (62054 bytes)Lesser Black-backed Gull graellsii NA1 adult, June 17 2002, Papegaaienbek Maasvlakte, the Netherlands.
Still no primaries shed by mid June.
adult LBBG in June, ringed in the Netherlands. (72840 bytes)Lesser Black-backed Gull graellsii PP4 7cy, June 02 2003, Dintelhaven Maasvlakte, the Netherlands.
Ringed as pullus June 28 1997, now 7cy. Present from 2001.
adult LBBG in June, ringed in the Netherlands. (62058 bytes)Lesser Black-backed Gull graellsii TV4 adult, June 18 2002, Dintelhaven Maasvlakte, the Netherlands.
Ringed as pullus at the Maasvlakte.
adult LBBG in June, ringed in the Netherlands. (56483 bytes)Lesser Black-backed Gull graellsii UX2 10cy, June 17 2002, Papegaaienbek Maasvlakte, the Netherlands. Ringed at Maasvlakte as pullus, June 21 1993.
1827er14.jpg (69521 bytes)Lesser Black-backed Gull graellsii ER14 8cy, June 02 2003, Dintelhaven Maasvlakte, the Netherlands. Ringed as pullus June 26 1996, now 8cy.
adult LBBG in June, ringed in the Netherlands. (90222 bytes)Lesser Black-backed Gull graellsii ES98 8cy, June 02 2003, Dintelhaven Maasvlakte, the Netherlands. Ringed as pullus June 26 1996, now 8cy.
ad LBBG in Januari-April, ringed in the Netherlands. (64361 bytes)Lesser Black-backed Gull graellsii ET55 21CY-22CY, April 2016 & June 2017, Leixões Harbour, Matosinhos, Portugal. Picture: José Marques.
adult LBBG in June, ringed in the Netherlands.Lesser Black-backed Gull graellsii EU06 6cy, June 16 2001, Dintelhaven Maasvlakte, the Netherlands. Female, with dark iris, mated with adult male right. Ringed as pullus June 28 1996.
adult LBBG in June, ringed in the Netherlands. (49236 bytes)Lesser Black-backed Gull graellsii EAS1 7cy, June 02 2003, Dintelhaven Maasvlakte, the Netherlands. Ringed as pullus June 24 1997, now 7cy.
adult michahellis in May, ringed in the Netherlands. (72689 bytes)Yellow-legged Gull michahellis CU9 13+cy & Lesser Black-backed Gull graellsii adult, Missouriweg - Maasvlakte, the Netherlands, May 12 2002 (51.59N,04.02E).
One of the few mixed pairs at Maasvlakte.
adult LBBG in June, ringed in the Netherlands.Lesser Black-backed Gull graellsii KC9 9cy, June 17 2001, Missouriweg Maasvlakte, the Netherlands.
Ringed July 07 1993.
adult LBBG in June, ringed in the Netherlands.Lesser Black-backed Gull graellsii KH0 9cy, June 26 2001, Bergen op Zoom, the Netherlands.
Ringed July 07 1993.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii N||A (was RV3), 19CY, June 06-14 2016, Maasvlakte, the Netherlands.
adult LBBG in June, ringed in the Netherlands. (41422 bytes)Lesser Black-backed Gull graellsii E053 9cy, June 02 2003, Missouriweg, Maasvlakte, the Netherlands.
Ringed as pullus June 01 1995, now 9cy.
adult LBBG in June, ringed in the Netherlands.Lesser Black-backed Gull graellsii E757 7cy, June 17 2001, Missouriweg Maasvlakte, the Netherlands.
Ringed July 09 1995.
adult LBBG in June, ringed in the Netherlands.Lesser Black-backed Gull graellsii E832 7cy, June 17 2001, Missouriweg Maasvlakte, the Netherlands.
Ringed July 09 1995.
adult LBBG in June, ringed in the Netherlands.Lesser Black-backed Gull graellsii EC13 adult, June 26 2001, Bergen op Zoom, the Netherlands.
adult LBBG in June, ringed in the Netherlands. (77018 bytes)Lesser Black-backed Gull graellsii EK85 8cy, June 02 2003, Missouriweg, Maasvlakte, the Netherlands.
Ringed as pullus July 09 1996, now 8cy.
adult LBBG in June, ringed in the Netherlands. (70789 bytes)Lesser Black-backed Gull graellsii EM30 8cy, June 02 2003, Missouriweg, Maasvlakte, the Netherlands.
Ringed as pullus July 10 1996, now 8cy.
adult LBBG in June, ringed in the Netherlands. (66296 bytes)Lesser Black-backed Gull graellsii EY87 6cy, June 17 2002, EMO - Maasvlakte, the Netherlands.
Ringed at Maasvlakte as pullus, July 03 1997.
adult LBBG in June, ringed in the Netherlands. (61343 bytes)Lesser Black-backed Gull graellsii PL3 6cy, June 24 2002, Papegaaienbek - Maasvlakte, the Netherlands.
Ringed at Maasvlakte as pullus, July 1997.
adult LBBG in June, ringed in the Netherlands. (70101 bytes)Lesser Black-backed Gull graellsii NLA 5.251.194 12CY, June 02 2003, Missouriweg, Maasvlakte, the Netherlands.
adult LBBG in June, ringed in the Netherlands. (61397 bytes)Lesser Black-backed Gull graellsii NLA 5.257.218 15cy, June 02 2003, Missouriweg, Maasvlakte, the Netherlands.
Ringed as pullus July 09 1989, now in 15cy with 3 pulli.
adult LBBG in June, ringed in the Netherlands. (57024 bytes)Lesser Black-backed Gull graellsii NLA 5.257.580 15cy, June 02 2003, Missouriweg, Maasvlakte, the Netherlands.
Ringed as pullus July 09 1989, now in 15cy.
adult LBBG in June, ringed in the Netherlands. (65264 bytes)Lesser Black-backed Gull graellsii NLA 5.280.505 10+CY, June 24 2002, Papegaaienbek - Maasvlakte, the Netherlands.
A dark individual.
adult graellsii in May, ringed in the Netherlands. (83502 bytes)Lesser Black-backed Gull graellsii NLA 5.417.298 adult, May & June 2015, Boulogne-sur-Mer, NW France. Picture: Jean-Michel Sauvage.
Ringed as breeding adult female.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii NLA 5.462.810 7CY, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii NLA 5.508.7xx adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii NLA 5.50x.555 adult, June 06 2016, Maasvlakte, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)Lesser Black-backed Gull graellsii NLA 5xxx adult, June 06 2016, Maasvlakte, the Netherlands.
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Denmark: intermedius.
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adult LBBG intermedius in September, ringed in Norway. (95609 bytes)Lesser Black-backed Gull intermedius VC5M adult, June & September 2010 & October 2012, Denmark, the Netherlands, Spain. Pictures: Kjeld Tommy Pedersen.
Ringed as breeding female.
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Spain: winter catches - no provenance on origin.
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3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)Lesser Black-backed Gull graellsii N0M2 adult, June 12 2016, Utrecht, the Netherlands.
Lesser Black-backed Gull graellsii ESI 61xxx01 adult, April & June 2017, Katwijk, the Netherlands.
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Arrested & Suspended Moult in June adult LBBG.
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suspended moult in adult LBBG in June. (76647 bytes)Lesser Black-backed Gull graellsii adult, June 02 2003, Maasvlakte, the Netherlands. Active moult in both wings, replacing P7 & P8.