third
calendar year: September
Last update of
this page: 10-10-05 06:19
In LBBG nearly all feathers
are being replaced now. Birds require fresh plumage with
adult-like upper-parts and under-parts. On average, coverts and tertials
are moulted into 3rd
generation feathers and mantle and scapulars become 4th
generation, after the moult to 3rd winter plumage.
From head to toe you can see
the following changes:
-
The bill will acquire a bill-band due to the hormonal changes
(3rd winter bill coloration).
There is quite some variation: some birds will get a
blackish base to the bill, while others only get a sharply defined
band across the gonys. The average birds (79 %) show an ”adult”
bill with a nice broad band over the upper and lower mandible. The
bill colours become paler, more
lemon and orange.
The legs turn pale pinkish yellow
or off-yellow.
-
The head becomes heavily streaked
as freshly moulted streaked
feathers grow in, replacing the white old, worn feathers. Some birds
acquire an almost greyish head.
-
The under-parts are
also moulted. Nearly 100% of the birds show
streaked under-parts, some having streaking in the hind-neck only.
-
Most 3cy LBBG
finish the moult of
mantle and scapulars (completed by mid September). The
upper-parts resemble adults now (being 4th or even 5th
generation).
-
Tertials
exist of
newly moulted 3rd generation feathers. They are moulted downwards,
giving a contrast between the new upper
tertials and the old lower tertials (with the central tertials
missing). The
difference is obvious with these new tertials adult-grey with a
white top, contrary the blackish centred old
tertials with black markings in the tip. Some birds show a more
blackish centre on new 3rd generation tertials. If so,
it's prominent in tertials #3-6.
At the end of September almost all 3cy have six new tertials, while some birds are still moulting the
final tertials.
- The wing-coverts have been moulted all to
3rd generation, but some 3cy LBBG have a moult-gap in the coverts or
still have some old coverts. 95 % follows the ordinary moult
sequence: first the inner 3-4 followed by the outermost feathers
(which are replaced inwards). Only 5% follows a random strategy and leave a few feathers in the greater coverts not replaced.
The under-wing coverts are replaced by
adult-like white feathers in a sequence similar to the upper-wing.
- The primaries are moulted
outwards and the primary-score at the end
of September averages over 40 (n = 153) with p6 full-grown on all bird,
as shown in the table below. The 3rd generation
primaries are very adult-like but p10 often shows an obvious smaller
mirror
-
The
secondaries are dropped inwards, starting next to the primaries.
In adults secondary moult starts from the moment
p6 is half way to fully grown. In 3cy, only a few
birds show secondary moult in September. The 3cy moult of secondaries is not related to moult of primaries. As shown
in the bottom table, birds with fully grown p8 or even p9
can still show complete old 2nd generation secondaries.
Such birds were found as late as October 10 2001.
-
The tail is new,
3rd generation in most cases (90%), but still in active moult
in some birds (5%) or complete 3rd generation, white or with very restricted black
markings (5%).
Old worn rectrices seen in September, without any black markings, are
probably 3rd generation as well, moulted early in
spring. Variation within 3cy and 4cy birds is huge and advanced 3cy
can be aged wrongly as delayed 4cy birds.
Remaining primaries
|
Full-grown primaries |
date: |
1/9 |
3/9 |
11/9 |
16/9 |
17/9 |
24/9 |
28/9 |
|
1/9 |
3/9 |
11/9 |
16/9 |
17/9 |
24/9 |
28/9 |
0 |
1 |
5 |
1 |
6 |
5 |
13 |
85 |
p10 |
|
|
|
|
|
|
|
p10 |
|
5 |
1 |
6 |
3 |
11 |
22 |
p9 |
|
|
|
|
|
3 |
16 |
p9 |
6 |
5 |
|
5 |
3 |
8 |
7 |
p8 |
1 |
2 |
|
4 |
1 |
11 |
63 |
p8 |
3 |
2 |
|
1 |
2 |
4 |
2 |
p7 |
2 |
9 |
2 |
10 |
8 |
10 |
30 |
p7 |
2 |
|
|
|
|
1 |
|
p6 |
4 |
4 |
|
4 |
4 |
13 |
7 |
|
|
|
|
|
|
|
|
p5 |
3 |
2 |
|
|
|
|
|
|
|
|
|
|
|
|
|
p4 |
2 |
|
|
|
|
|
|
lesser
coverts |
greater
coverts |
week
no: |
1/9 |
3/9 |
11/9 |
16/9 |
17/9 |
24/9 |
|
|
1/9 |
3/9 |
11/9 |
16/9 |
17/9 |
24/9 |
|
0 |
|
|
|
|
|
|
|
0 |
1 |
|
|
|
|
|
|
1 |
2 |
2 |
|
|
|
|
|
1 |
|
4 |
|
|
|
|
|
2 |
1 |
2 |
|
|
|
|
|
2 |
3 |
1 |
|
1 |
|
|
|
3 |
3 |
4 |
|
|
|
|
|
3 |
2 |
2 |
|
1 |
|
|
|
4 |
4 |
3 |
1 |
4 |
2 |
3 |
|
4 |
2 |
4 |
|
3 |
2 |
1 |
|
5 |
2 |
6 |
1 |
14 |
11 |
34 |
|
5 |
4 |
6 |
2 |
13 |
11 |
36 |
|
|
New
Tertials |
|
1/9 |
3/9 |
11/9 |
16/9 |
17/9 |
24/9 |
|
0 |
|
|
|
|
|
|
|
1 |
|
1 |
|
|
|
|
|
2 |
|
|
|
|
|
|
|
3 |
1 |
2 |
|
|
|
|
|
4 |
4 |
3 |
|
2 |
|
|
|
5 |
3 |
3 |
|
2 |
|
2 |
|
6 |
4 |
8 |
2 |
14 |
13 |
35 |
|
n: |
12 |
17 |
2 |
18 |
13 |
37 |
|
Coverts score:
0 = all coverts old, 1 = 25% new, 2 = 50% new, 3 = 75% new, 4
= 99% new, 5 = 100% new
|
primary
moult score in twenty-two 3cy LBBG, Boulogne-sur-Mer, NW
France, September 21 2002. |
|
p8
old |
p9
old |
p10
old |
no
old |
p8
new |
|
|
|
4 |
p7
new |
|
3 |
2 |
9 |
p6
new |
1 |
2 |
|
|
p5
new |
1 |
|
|
|
|
3cy
LBBG:
Relation between
secondary
moult, new
primaries and date.
|
|
all
S old |
outer
miss, inner old |
outer
grow, inner miss |
outer
new, inner grow |
all
S new |
S
not seen |
p4 |
4,2 |
|
|
|
|
|
p5 |
6,3 |
|
1 |
|
|
|
p6 |
1,3,2 |
5,1,2,2 |
3,6 |
4 |
|
2,1 |
p7 |
1,1 |
1,2,1 |
1,1,1,2,2 |
3 |
4,2,1,3,2 |
1,2 |
p8 |
|
|
1 |
2 |
1,2,1,6 |
2 |
p9 |
|
1 |
|
|
2 |
|
p10 |
|
|
|
|
1 |
|
all S old:
all secondaries old, i.e. still 2nd generation.
outer miss, inner old: outer secondaries shed (S1-S9),
while inners are still old (S10~S18).
outer grow, inner miss: S1-S9 are growing, while
S10~S18 are shed.
outer new, inner grow: S1-S9 are new 3rd
generation, S10~S18 are growing.
all S new: all secondaries are new 3rd
generation.
S not seen: secondary moult stage not certain.
p4,...p10: most recent moulted full-grown primary.
date:
28/08/2001, 01/09/2001, 11/09/2001,
16/09/2001, 17/09/2001,
24/09/2001 |
|
|
LBBG A18
3cy, September 18 2004,
IJmuiden, the Netherlands.
LBBG intermedius J1E7
3cy, September 01 2003,
Naaktstrand, Maasvlakte, the
Netherlands.
LBBG intermedius NOS
4217224 (3cy) September
02 2003, Westkapelle, the
Netherlands. Arrested moult at P7.
LBBG intermedius SVS
8103331 3cy, September
02 2003, Westkapelle, the
Netherlands.
Photo
1: LBBG 3cy, September 02 2001, Missouriweg Maasvlakte, the
Netherlands.
Photo
2: LBBG 3cy, September 02 2001, ECT parking Maasvlakte, the
Netherlands.
Photo
3: LBBG 3cy, September 02 2001, ECT parking Maasvlakte, the
Netherlands.
Photo
4: LBBG 3cy, September 17 2001, Vuurtorenvlakte Maasvlakte, the
Netherlands.
Photo
5: LBBG 3cy, September 30 2001, Dannes - Le Portel, France.
Photo
6: LBBG sub-adult, September 24 2001, IJmuiden, the Netherlands.
Photo
7: LBBG 3cy, September 24 2001, IJmuiden, the Netherlands.
Photo 8: LBBG 3cy, September
16 2002, Scheveningen, the Netherlands.
Photo 9: LBBG 3cy, September
22 2002, Le Portel, France.
Photo 10: LBBG 3cy, September
22 2002, Le Portel, France.
|