General
description:
By August, the number of migrating
Herring Gull and Lesser Black-backed Gull increase dramatically in
NW France, but 3cy Yellow-legged Gull can still be found commonly in
the vicinity of Etaples and Boulogne-sur-Mer (see Le
Portel).
The general appearance of 3cy michahellis is rather
straightforward. Most birds can be recognized by the plain grey
upper-parts and coverts, combined with some retained barred
wing-coverts, especially in the carpal edge. The bare parts are
typically a red orbital
ring, a black band on the bill and yellow legs. An example of the average
August 3cy michahellis
can be found in the sixth image, an individual
with adult-like upper-parts, some barred wing-coverts in the carpal
edge, some new white tail-feathers and the new primaries just
visible beneath the tertials.
In the surveys of the last days of
July 2001 3cy michahellis showed an average primary moult score of
8.5 remaining old primaries. By the last week of August, 3cy michahellis
may show two, one or no old primaries. The table below shows figures
for July and September.
For
details on differences between michahellis and cachinnans, see
e.g. the article by R. Klein & D. Gruber in Limicola, April
1997.
Moult
stage and pace of remiges and rectrices:
The complete moult started by mid-May,
when 3cy birds dropped the innermost primary P1. By early July, the
average score has strongly increased: P7-P10 are still old second
generation and the inner primaries are new. Since the
tips of P5 and P6 normally extend beyond the tertials (especially
the abraded July tertials), the moult gap of missing P5 and P6 is
obvious.
3cy michahellis continue shedding primaries throughout the
month, and by the end of July, the average primary moult score for
remaining primaries is 8.5. With this score, 3cy birds are about two primaries behind compared to
2cy birds
(which have a score of 9.2: 45% of 2cy have shed all primaries by
late July).
As in adults, the gap between full-grown primaries and remaining old
primaries is in general about three feathers, or slightly more as the table in the July
section shows. This large moult gap is
typical for michahellis, and not found in e.g. Dutch argenteus.
By early August, 3cy michahellis has an average score P6
fully grown, so the majority of resting birds shows new
primaries in the folded wing. This score increases gradually
during the month. We have no late-August or early-September scores,
but by the third decade of September the primary moult score
exceeded 8.0. The score for the end of August must be about P7.
By the time P4 is fully grown, 2cy
birds normally start the complete moult in secondaries and rectrices.
The ordinary moult strategy in 3cy birds may be slightly later
(starting when P5 is fully grown. Note however that individual
variation may exist, particularly in 3cy michahellis
rectrices moult as this moult may follow an irregular pattern in
earlier months. The
partial spring moult has finished in June and a small minority of 3cy
June michahellis
have included some rectrices in this partial moult. Hence, it's not uncommon
to find 3cy birds showing plain white tail-feathers, in some birds
creating a 'blocked' tail pattern already, prior to the complete
moult.
On average, the common sequence is centrifugal moult:
starting with the inner tail-feathers and continuing symmetrically
outwards to both outer feathers. The moult in the rectrices is still
in progress by late-August and the new new third generation
tail-feathers normally show some black markings.
Secondary moult starts in early August. The images on the July page
showing
birds with open wings illustrate that all outer secondaries are
still present. By early August, the outermost secondaries (S1 &
S2, at the division between secondaries and primaries) are normally
dropped individually, but the central secondaries are shed almost as
in small groups, creating large gaps of missing secondaries, nicely
illustrated by this flying 2cy
bird and this stretching 3cy
August michahellis.
The innermost secondaries are still second generation. Since many
birds moult the inner greater coverts, the secondaries underneath
are exposed, nicely demonstrated by this
bird.
Timing and
strategy of body & covert moult in August
Wing-covert moult in 3cy and sub-adult michahellis vary
between individuals. See e.g. 365D,
showing extensive moult in the wing-coverts (the sequence is mentioned in the accompanying
text). Such birds contrast strongly with e.g. this
bird, which has almost all the wing-coverts old.
In general, conclusions based on average 3cy michahellis
are:
- Moult of head feathers is in
progress, but almost finished in advanced individuals, creating a streaked head.
See here (streaking around the
eye) and here
(streaking around the eye and in the hind-neck).
- Average 3cy birds show plain adult-like
grey scapulars from July onwards. It's
difficult to score missing feathers in the complete grey
upper-parts, hence we have no scapular moult scores. In general,
the scapulars in the lowest row look very fresh, without
abrasion at the fringes and lacking the faded grey tone, so
scapular moult may be near its end in 3cy michahellis by
the end of July and early August. The odd individual may still
be found missing rear lower scapulars, normally the once moulted
last in line.
In some
individuals, the scapular region contains some old and abraded feathers
(see here, in the lowest scapulars),
sometimes showing a brown hue, others are bleached white-grey
with only a slightly darker shaft-streak. However, most birds have the
entire back adult-like grey.
-
The complete wing-covert moult is in progress throughout July
and August. The
ordinary moult strategy and sequence start in the outer median
coverts, the innermost medians and innermost greater coverts, the upper tertial
and the outer lower lesser coverts. By August, the median and
lower lesser coverts have been replaced and 3cy birds are
missing the outer greater coverts and some lesser coverts. By
the end of August, the outer greater coverts have been replaced
as well and the gap in the greater coverts will be closed at
greater coverts #7-#8 in birds following the ordinary moult
strategy, see e.g. here
(missing inner greaters) and here
(closing the gap at #7-#8).
By late-August, the most advanced birds have replaced all the
wing-coverts and may start again a new additional cycles of
wing-covert moult: the partial wing-covert moult in autumn. They
normally start at the oldest wing-coverts, i.e. the median
coverts and lower lesser coverts. See e.g. here.
- In the tertials, the upper 3-4
feathers have been replaced by the end of August and the lower
two tertials are missing. During the month of September, 3cy michahellis
moult the complete secondary row and tertials (which actually
belong to the secondaries). The gap in the secondaries will be
closed at about #18.
See Topography
Section for explanation of feather tracts.
Tables:
The surveys
were done at resting and preening 3cy sub-adults (with at least
no white mirror at P10 and at least some obvious second summer feathers).
Primary
moult score of 3cy michahellis Yellow-legged Gull, at
Etaples, NW France (50.43N,
01.37E) on August 09 2003. |
. |
P5
fg |
P6
fg |
P7
fg |
?
fg |
n: |
P8-P10 |
3 |
- |
- |
1 |
4 |
P9-P10 |
1 |
8 |
- |
1 |
10 |
P10
old |
- |
5 |
- |
- |
5 |
no
old P |
- |
- |
3 |
- |
3 |
n: |
4 |
13 |
3 |
2 |
22 |
n
= 22; m old P = 1.7; SD old P = 0.95.
Group
of 328 michahellis present: 28 juveniles, 37 2cy, 35 3cy, 218
(sub)adults.
P8-P10 = P8-P10 are still old and present.
? fg = fully grown not known. Fully grown primary score most
probably P6 or less.
fg = fully grown. In our field surveys, a primary is 'fully
grown' when it exceeds the previous feather in the folded
wing. Actually, such 'fully grown' feathers may have to
grow a little, but under field circumstances, it's very
difficult to find out if the waxy sheath, which is the
standard measurement for fully grown primaries, is still
present. |
Primary
moult score of 3cy michahellis Yellow-legged Gull, at Etaples,
NW France (50.43N,
01.37E) on August 11 2003. |
. |
P5
fg |
P6
fg |
P7
fg |
P8
fg |
n: |
P8-P10 |
5 |
- |
- |
- |
5 |
P9-P10 |
3 |
24 |
- |
- |
27 |
P10
old |
- |
10 |
6 |
- |
16 |
no
old P |
- |
- |
2 |
- |
2 |
n: |
8 |
34 |
8 |
- |
50 |
n
= 50; m new P = 6.0; SD new P = 0.57; m old P = 1.7; SD old P =
0.70.
Further comments / notes: see previous table. |
3cy
michahellis primary moult scores: Boulogne-sur-Mer, NW France, July
& September 2001 & 2002. |
new
primaries |
July
29 2001
Etaples |
Sept
22 2002
Etaples |
Sept
29 2001
Etaples |
Sept
29 2001
Le Portel |
p5 |
1 |
- |
- |
- |
p6 |
12 |
- |
- |
- |
p7 |
25 |
- |
- |
- |
p8 |
4 |
33 |
2 |
2 |
p9 |
- |
7 |
23 |
16 |
p10 |
- |
- |
8 |
5 |
n: |
42 |
40 |
33 |
23 |
m: |
6.8 |
8.2 |
9.2 |
9.2 |
s: |
0.66 |
0.38 |
0.53 |
0.53 |
notes:
3cy birds selected on bare part coloration, tail pattern, pattern on
inner secondaries and outer greater primary coverts. Scores of
longest fully grown new third generation primaries.
|
|
Photo
7928: Michahellis 3cy, August 15 2003, Westkapelle, the Netherlands
(51.33N,
03.25E).
Compare grey tone to Lesser Black-backed Gull.
|