| General
            description: Early July and from
      July 26-30 2001 we did small surveys on Yellow-legged Gulls at Le
      Portel
      (near Boulogne-sur-Mer) and Etaples (near Le Tourquet). In these surveys, we
      scored moult stages to determine timing of moult in primaries (and wing-
      coverts and body-feathers) in sub-adult michahellis.By June, sub-adult michahellis
            occurs only in small numbers at Le Portel and the near surrounding.
            But by July, more Yellow-legged Gulls disperse to northern quarters
            to rich feeding grounds and to moult the primaries. At the dumping site of Dannes,
            near Le Portel, mixed flocks of mainly Lesser Black-backed
            Gulls (100's) from NW and N Europe can be found including tens of
            adult and sub-adult Yellow-legged Gulls.
 Regarding identification: the difference between 3cy birds and 4cy
            birds can best be established by the outer primary, which is still
            unshed by July: 4cy birds show a small but obvious white mirror on
            P10, while 3cy birds have the (second generation) outer primary all
            brown. In birds with very worn outer primary tips it may be
            difficult to ascertain whether a mirror has been present or not. Other
            clues can be found in the bare part coloration, with 4cy strongly
            resembling adults. Complete
            moult in July: As can be seen in most of the images,
            sub-adult Yellow-legged Gulls look very untidy and mottled in their
            July plumage. Simultaneous moult is in progress in the primaries,
            wing-coverts, tertials, scapulars and the
            tail-feathers. Fresh moulted wing-coverts can be found in
            the median row and outer lower lesser coverts in most cases and these fresh grey median coverts
            normally strongly contrast with the brown-hued lesser coverts and
            old greater coverts. The old coverts have very abraded fringes and
            often a brown hue. By early July, the
            upper tertial has been shed in many birds, as is the case with the
            central tail-feathers.
             Moult
            stage and pace of remiges and rectrices: The table below shows the average
            score of immature Yellow-legged Gulls, range between 3cy birds to
            5cy birds. Individuals in 3cy show no mirror on P10, 4cy birds have
            various wing-coverts, tertials or tail-feathers with black markings
            and 5cy birds are very adult-like. All birds showing black on the
            bill combined with dark outer primary coverts were included in this score.
            It's hard to predict the score of new primaries; too many resting
            and sleeping birds could not be scored on the new fresh grown inner
            primaries (over 35% of the immature birds present were not scored in
            detail and are clustered in the table under "P?"). The
            average score of old primaries is 8.1, hence on average less than
            two old primaries are still present in the last week of July. In 3cy birds, the early-July score
            averages 7.0 (n=33 on July 10 2002), so three old primaries left. By the end of July, the average primary moult score is
            8.5 (n=39 on July 29 2001; n=53 on July 30 2001), only slightly
            higher than 4cy michahellis. In adult birds, the early-July
            score
            averages 6.3 (n=66 on July 11 2001), so more than three old
            primaries left in most birds. By the end of July, the average primary moult score is
            7.6 (n=50 on July 28 2001) and 7.7 (n=56 on July 29 2001) in adults,
            so slightly behind compared to 4cy michahellis. Adult birds
            were selected on plain grey primary coverts. By the time P4 is fully grown, 2cy
            birds normally start the complete moult in secondaries and rectrices. 3cy birds
            start slightly later, when P5 is fully grown, although individual
            variation may exist. Sub-adult michahellis probably follow a
            moult timing similar to 3cy birds. Old tail-feathers may still show
            much black, typically found on third generation rectrices, but old
            feathers are not necessarily third generation in michahellis.
            As can be seen in previous sections, tail-feathers may be included
            in the partial autumn moult in both 1cy and 2cy, so old
            tail-feathers in July sub-adults may be completely white as well.
            However, the new white tail-feathers can be recognized by the neat
            fringes and tips. Secondary moult hasn't started yet in late-July, but
            is initiated in early-August.
 Timing and
            strategy of body & covert moult in July Wing-covert moult in 3cy and sub-adult michahellis vary
      between individuals. In general, conclusions based on average sub-adult michahellis
      are: 
              Moult of head feathers is in
                progress, creating a "moth-eaten" appearance.sub-adult birds show plain adult-like
      grey scapulars. It's
                difficult to score missing feathers in the complete grey
                upper-parts, hence we have no scapular moult scores. In general,
                the upper lower scapulars look very fresh, the lower upper
                scapulars look older and worn and some of the scapulars in the
                lowest row seem to be missing in sub-adult michahellis by
                the end of July.
                The complete wing-covert moult is in progress throughout July. The
                ordinary moult strategy and sequence start in the outer median
                coverts, the innermost medians and innermost greater coverts, the upper tertial
                and the outer lower lesser coverts. Early July, the new median
                coverts are visible (first the outer median coverts), more so as
                the outer lower lesser coverts are dropped. By the end of July,
                the complete median covert bar is renewed, the lower lesser
                coverts are growing, the outer greater coverts are missing
                (exposing the secondaries underneath) and the innermost greater
                coverts have been renewed. In the tertials, the new upper two
                feathers are visible and the central tertials are dropped. See Topography
      Section for explanation of feather tracts.
             Tables: 
              
            
              
                | sub-adult
                  michahellis
                  (3cy-5cy) remaining old and fully grown new primaries, July 28
                  2001 Etaples. |  
                |  | P? | P4
                  new | P5
                  new | P6
                  new | P7
                  new | n
                  = |  
                | P6
                  old | 3 |  |  |  |  | 3 |  
                | P7
                  old | 10* | 3 | 2 |  |  | 15 |  
                | P8
                  old | 22 | 17 | 14 | 2 |  | 55 |  
                | P9
                  old |  |  | 18 | 6 | 1 | 25 |  
                | P10
                  old |  |  |  | 1 |  | 1 |  
                | no
                  old | 1 |  |  |  |  | 1 |  
                | n
                  (total): 100m-old (average): 1.9 remaining old primaries; so moult score at
                  8.1
 SD = 0.77
 P?: many birds sleeping, fully grown primaries hidden by
                  tertials, so P4 or less.
                  
      Aggregated survey done at resting and preening sub-adults
                  in 3cy, 4cy and 5cy (with at least black central markings on
                  the outer primary coverts).
 *: including one bird with P2 fully grown.
 |    |  Photo
    01: michahellis   sub-adult, July 10 2002, Etaples, NW France. 
    A sub-adult with active moult in the
    tail-feathers and some black on the cutting edge of the lower mandible. |