General
description:
By June the primary moult is obvious as
the outer flight-feathers are dropped now. Most birds look shabby, with
obvious gaps in
the median and greater wing-covert rows. The feathers which were included in the
post-juvenile moult are normally left out of this spring moult until July. The exact figures are hard to obtain, as birds
which do drop these earlier moulted feathers are not recognized of course
(unless birds are ringed and reported several times). However,
irregular moult sequences are not found in the tertials, as in most birds replacement
in the tertials follows the ordinary sequence, from tertial #1 downwards.
The majority of 2cy michahellis is easily identified by (some)
adult-like grey
scapulars, darker grey than in Herring Gull, almost blue-grey, and different from the
silvery grey third generation feathers in argenteus. Check out the
2cy argenteus images in the May
Section and June Section.
By June, 2cy michahellis
in complete moult (second pre-basic moult) can be found commonly around
the harbour of Etaples in NW France with a few 10's of birds in the
direct surrounding of the fish-market and many immature birds using the
near landfill of Dannes to feed on.
Identification:
The general appearance of 2cy michahellis is rather different from 2cy
June Herring Gulls from Belgium,
the Netherlands,
especially regarding the pattern on fresh scapulars and wing-coverts.
However, some local Herring Gulls from NW France may look very similar.
have a look at these ringed Herring Gulls: this,
this and this
one.
Strong and powerful male michahellis are readily easy recognized
between 2cy argenteus. Examples of average May michahellis
can be found in photo 4928 and in photo
1734. Those powerful birds (presumable males) show a combination of
almost completely white under-parts, some grey adult-like
scapulars, extensive moult in the wing-coverts and by June, an average
moult score of 6.0 (P6 has been dropped, see table below). Such features are indicative for
southern breeding taxa, as michahellis is, but doesn't necessarily
exclude French Herring Gulls. Michahellis moult
timing may show
overlap with argenteus from southernmost populations and local argenteus
(argenteus
breeds in the industrial area of Boulogne/Mer).
Some Lesser Black-backed
Gulls graellsii and intermedius may resemble miniature michahellis,
as this taxon may show advanced moult in wing-coverts and tertials as
well. However, ID is often straightforward as its mainly the smaller taxon intermedius
showing most advanced moult in LBBG. Furthermore, LBBG has a much broader
tail-band, darker inner primaries and the last required scapulars often
show much darker grey centres. Check out the 2cy June LBBG pages here.
Remember that many LBBG winter far south (NW Africa), suspending moult
during migration in June, thus showing only small
moult gaps, contra extensive moult in June michahellis,
showing obvious large moult gaps in the
wing-covert rows.
Remember that our surveys took place in Etaples (location information on the Le
Portel Page), where you find michahellis from the western Mediterranean:
mainly from S
France, a few from Italy, Switzerland and NE Spain.
Grey-based
scapulars or barred scapulars:
Images 1734 and 4928 represent strong male michahellis,
easily recognized by the jizz, and showing barred wing-coverts
and scapulars. The base colour is often brown-buffish in fresh feathers (by
May - June) and the pattern, which
can be bold and obvious on the scapulars, is normally repeated on the wing-coverts and
result in a complete barred plumage in freshly moulted birds.
Some female michahellis may show barred scapulars as well, but at
Etaples, birds showing extensive grey scapulars are often smaller
birds, so presumed females, as this bird demonstrates.
The causality is one-way: smaller birds often show grey scapulars, but
large birds do not necessarily have to show barred scapulars, as this
bird and this bird show.
Some michahellis may appear exceptionally advance
in moult, as the female in image 1357
shows. This bird already show plain grey wing-coverts in the medians as
well. has Probably, previous to the complete moult in summer, this individual has
replaced many wing-coverts and tertials to second generation feathers. By
June these coverts aremoulted again and may appear adult-like. Remember
that birds with advanced post-juvenile moult do not necessarily
develop all-grey scapulars in an early stage but may grown in fresh barred
feathers as well. Therefore, by the barred
patterns, these advanced birds do not necessarily look or appear more adult-like than
others, as this bird demonstrates: all
wing-coverts have been replaced, but they all appear immature. We don't
know whether development of plain grey third generation scapulars or barred
third generation scapulars is gender related, but on average, it seems the
smaller birds (presumable females) develop more often grey scapulars than
barred.
Moult stage and
pace:
What's the average moult in June? We
described 24 2cy birds in detail on two visits at Etaples,
Calais region and the following pattern can be
extracted:
By mid-June, primary moult is now visible
in the folded wing. The average primary moult score: P6 missing and P7-P10 the last juvenile
primaries. Most
advanced birds by mid-June show an obvious gap in the central primaries,
already missing P7 (almost 20% of the birds). Out of 41 birds, none had a
score below "P3 missing", a figure matched by the approximately
200 Herring Gulls L. argentatus s.l. from the same location and
from the Netherlands (one week later). The average moult
stage of L. argentatus from this location almost equals michahellis,
with bulk of the
present birds missing P5 or P6 (table below). Interestingly, the primary
moult score of local argenteus is slightly higher than argentatus
s.l. average, as preliminary figures of metal-ringed French argenteus
show.
The majority (about 75%) of 2cy michahellis
have all the rectrices still juvenile. From April onwards, moult in the
rectrices is still postponed but next month, when the inner 4 primaries
are grown, the moult in both secondaries and rectrices will start.
In several sections of 1cy michahellis
it's mentioned that birds include wing-coverts in the post-juvenile moult
(unlike argentatus / argenteus where this post-juvenile
moult is confined to the scapulars). To distinguish between 2cy birds
which have still suspended moult (after the post-juvenile moult) and birds
which have really started to replace wing-coverts, we separate between old
second generation feathers and fresh second generation feathers in the
table below. This is much easier said than done as the old second
generation feathers may have been required as early as late-August and
these wing-coverts may be extremely similar to the abraded juvenile
wing-coverts. Close views under excellent light conditions are necessary
to distinguish between the two by June, see e.g. this
individual.
Except the difference in timing of moult, these new second
generation feathers may show a different pattern as well, with fresh
second generation feathers moulted from April and May onwards looking 'less
immature': the base colour of the feathers is not buff-brown but
buff-grey or feathers may be "grey-based" and the pattern may be less well-defined.
Its interesting to see that the
wing-coverts
which were included in the post-juvenile moult are left out of the first
stage of the complete moult in early summer (until June at least) and are
replaced in a later stage. This is nicely demonstrated by a Swiss
metal-ringed michahellis Sempach 885629. The exact figures
of such moult sequences are hard to obtain, as birds
which do drop these earlier moulted feathers are not recognized of course. However,
irregular moult sequences are not found in the tertials, as in most birds replacement
in the tertials follows the ordinary sequence, from tertial #1 downwards.
The table below present figures of active wing-covert moult by mid-June.
The progress in wing-covert moult is obvious with moult gaps in almost all
wing-covert tracts, unless all coverts in arow have been replaced
completely. Leaving older second generation median coverts in place, while
moult progresses in other tracts is a very common phenomenon, as 9 of 14
birds demonstrate. This is even more clear in the lower lesser coverts,
where the figure is 7 out of 7. A few individuals have completely finished
moult in the greater coverts, but all birds present are actively moulting
lesser coverts. See Gull
Topography for explanations of feather tracts.
As can be seen in many
images of May, June and July 2cy michahellis, the sequence of moult
in the lower lesser coverts is sometimes hard to obtain correctly.
Post-juvenile feathers are maintained over a longer period and result in a
'jumpy' moult pattern and only after these old post-juvenile feathers are
replaced for new third generation feathers, the lower lesser covert row
appears neat and tidy. It's the end of July then...
Moult to third
generation scapulars may start as early as mid-January in 2cy michahellis.
By June, all present birds show active moult in the scapulars, with
largest numbers for the class 75%-99% third generation scapulars (8
birds). 50% of the birds (n=24) have over 50% of the
visible area (optical measurement) of the scapulars renewed. Previous sections of 2cy michahellis
show the most commonly seen moult sequence in the scapulars, starting with
the lower upper scapulars and finishing with the rear lowest scapulars. So
not surprisingly, its mainly the lower scapulars which have been
replaced by May, as is illustrated by this bird. Nevertheless,
some birds may still moult upper scapulars. Some 2cy michahellis
showed a red orbital ring by mid-April to mid-May. This coloration might be initiated
by hormones in early spring. By June and July, the red coloration has
weakened and the orbital ring looks immature again.
Tables:
remaining
old primaries in 2cy Larus michahellis, Etaples, NW France on June
13 & 14 2002. |
P5-P10 |
P6-P10 |
P7-P10 |
P8-P10 |
m: |
n: |
SD: |
1 |
7 |
25 |
8 |
4.0 |
41 |
0.69 |
Primary
moult score in 2cy Larus michahellis, Etaples, NW France on
June 26 2003. |
. |
P2
fg |
P3
fg |
P4
fg |
P5
fg |
P6
fg |
?
fg |
n: |
P5-P10 |
1 |
- |
- |
- |
- |
- |
1 |
P6-P10 |
- |
- |
- |
- |
- |
- |
- |
P7-P10 |
- |
2 |
3 |
- |
- |
1 |
6 |
P8-P10 |
- |
- |
2 |
13 |
- |
- |
15 |
P9-P10 |
- |
- |
- |
6 |
- |
- |
6 |
n: |
1 |
2 |
5 |
19 |
- |
1 |
28 |
n
= 28; m old P = 3.1; SD old P = 0.88.
P6-P10 =
P6-P10 are still old and present.
? fg = fully grown not known. Often referring to sleeping birds with
folded wings, so fully grown primary score most probably P6 or less.
fg = fully grown. In our field surveys, a primary is 'fully grown'
when it exceeds the previous feather in the folded wing. Actually,
such 'fully grown' feathers may have to grow a little, but under
field circumstances, it's very difficult to find out if the waxy
sheath, which is the standard measurement for fully grown primaries,
is still present. |
Primary
moult score in 2cy Larus michahellis, Etaples, NW France on
June 27 2003 (17:30 h). |
. |
P3
fg |
P4
fg |
P5
fg |
P6
fg |
?
fg |
n: |
P6-P10 |
- |
- |
- |
- |
- |
- |
P7-P10 |
- |
1 |
- |
- |
4* |
5 |
P8-P10 |
- |
- |
13 |
- |
7 |
20 |
P9-P10 |
- |
- |
16 |
3 |
- |
19 |
P10
old |
- |
- |
- |
1 |
- |
1 |
n: |
- |
1 |
29 |
4 |
11 |
45 |
n
= 45; m old P = 2.6; SD old P = 0.71.
Group: 4x 2cy michahellis not scored; total michahellis
present: 185.
* = including metal ring. |
Primary
moult score in 2cy Larus michahellis, Etaples, NW France on
June 28 2003 (08:30 h, low tide). |
. |
P5
fg |
P6
fg |
?
fg |
n: |
P7-P10 |
1 |
- |
4 |
5 |
P8-P10 |
44 |
3 |
11 |
58 |
P9-P10 |
11 |
18 |
3 |
32 |
P10
old |
- |
1 |
- |
1 |
n: |
56 |
22 |
18 |
96 |
n
= 96; m old P = 2.7; SD old P = 0.58.
Group: 4x 2cy michahellis not scored; total michahellis
present: 300+. |
primary
moult
scores in 2cy Larus argentatus s.l. (most argenteus):
a) Etaples, NW France on June 13 2002.
b) Scheveningen, the Netherlands on June 21 2002. |
|
p4
miss |
p5
miss |
p6
miss |
p7
miss |
m: |
n: |
SD: |
a |
7 |
23 |
56 |
10 |
5.7 |
96 |
0.75 |
b |
9 |
36 |
43 |
13 |
5.6 |
101 |
0.83 |
rectrices
moult
scores in 2cy Larus michahellis, Etaples, NW France, June 13
& 14 2002. n =
22
|
all
juvenile |
>
0 new rectrices |
17 |
5 |
moult
scores in several feather tracts of 2cy Larus michahellis,
June 13 & 14 2002, Etaples, NW France. Total n = 41.
Partial n fluctuates due to observation circumstances. |
tertials |
>2
juvenile |
<3
juvenile, active moult |
all
2nd generation |
n
= 24 |
19 |
4 |
1 |
greater
coverts |
juvenile
+ missing |
juv
+ some post-juv + active moult |
juvenile
+ fresh 2nd gen. |
all
new 2nd gen. |
n
= 23 |
7 |
6 |
8 |
2 |
median
coverts |
juv
+ missing |
active
moult: some fresh |
complete
post-juv + new 2nd gen. |
complete
new 2nd gen. |
n
= 23 |
1 |
8 |
9 |
5 |
lower
lesser coverts |
juv
+ missing |
active
moult: some fresh |
complete
post-juv + new 2nd gen. |
complete
new 2nd gen. |
n
= 23 |
2 |
14 |
7 |
- |
lesser
coverts |
most
juv + < 50% missing |
>
50% missing |
most
2nd gen. + < 50% missing |
n
= 21 |
5 |
13 |
3 |
scapulars |
<
25% 3rd gen. |
25%
< 3rd gen. < 50% |
50%
< 3rd gen. < 75% |
75%
< 3rd gen. < 100% |
n
= 24 |
7 |
5 |
4 |
8 |
|
171Z michahellis 2cy,
June 27 2003, Etaples, France. A ringed michahellis
from S France, missing one leg.
|