ORG SITE
YLG
1cy May
YLG
1cy June
YLG
1cy July
YLG
1cy August
YLG
1cy September
YLG
1cy October
YLG
1cy November
YLG
1cy December
YLG
2cy January
YLG
2cy February
YLG
2cy March
YLG
2cy April
YLG
2cy May
YLG
2cy June
YLG
2cy July
YLG
2cy August
YLG
2cy September
YLG
2cy October
YLG
2cy November
YLG
2cy December
YLG
3cy January
YLG 3cy February
YLG 3cy March
YLG 3cy April
YLG 3cy May
YLG 3cy June
YLG 3cy July
YLG 3cy August
YLG 3cy September
YLG 3cy October
YLG 3cy November
YLG 3cy December
YLG
sub-ad Jan.
YLG sub-ad Feb.
YLG sub-ad March
YLG sub-ad April
YLG sub-ad May
YLG sub-ad June
YLG sub-ad July
YLG sub-ad Aug.
YLG sub-ad Sept.
YLG sub-ad Oct.
YLG sub-ad Nov.
YLG sub-ad Dec.
YLG
adult January
YLG adult February
YLG adult March
YLG adult April
YLG adult May
YLG adult June
YLG adult July
YLG adult August
YLG adult September
YLG adult October
YLG adult November
YLG adult December
|
This
website deals with the Yellow-legged Gull taxon michahellis, which
is a common migrant from July to December in NW Europe. After extensive expansion
of the breeding population during the last three decades, it nowadays can
be found breeding in Belgium, the Netherlands and Britain in mixed couples
with both Herring Gull (argenteus) and Lesser Black-backed Gull (graellsii).
There are subtle differences between the populations from the
Mediterranean, Atlantic coast of Portugal and Morocco and from the islands
in the Atlantic. Most pronounced differences can be found in the taxon atlantis,
now regarded as full species by some authors.
L. michahellis: adult July
General
description:
Early July and from July 26-30 2001 we
did small surveys on Yellow-legged Gulls at Le
Portel (near Boulogne-sur-Mer) and Etaples (near Le Tourquet). In
these surveys, we scored moult stages to determine timing of moult in
primaries (and wing- coverts and body-feathers) in adult michahellis.
Adult michahellis is readily easy identified by July, as only
limited adult Herring and Lesser Black-backed Gulls are present.
For details on differences between michahellis and cachinnans, see
e.g. the article by R. Klein & D. Gruber in
Limicola, April 1997.
Moult of remiges and rectrices in
July:
The tables below show
the moult progress in a two week interval at Etaples, near
Boulogne-sur-Mer, NW France (click here to find maps of locations and pictures of Le Portel surroundings). In the
second week of July, the adult michahellis primary score is
slightly over 6: 6.3 in the first table. By the end July, primary moult
continues, only leaving the outer four or five primaries unshed. It's hard
to draw any conclusions about the moult stage of new grown primaries in
July, as in most birds the exact primary moult score could not be
established. However, the score of remaining old primaries was established
at 2.4, with n = 106, weight SD = 0.63, July 28-29 2001 at Etaples. The
very narrow range of moult scores (expressed by the low SD value) is one
of the most striking characteristics when scanning a group of michahellis.
The spread around the mean value is often near perfect and in this respect
differs strongly from e.g. Lesser Black-backed Gull, which has
high SD values, in many cases caused by a mix of different populations.
For more details on LBBGs, click
here. A gap of three primaries is common in adults, as can be
found in the tables as well; e.g. P4 fully grown, P5 almost fully grown,
P6 almost at half of the final length, P7 just shed and P8 still old.
The secondary moult is just not started by the end of July. In immatures
the secondary score is slightly advanced compared to adults and by the end
of July, third calendar year michahellis drops the 2nd,
3rd and 4th secondaries (the outermost secondaries).
Near adults (4cy birds) are similar in moult pace to the adults and
normally show a complete secondary row. The same is true for the rectrices. Normally, adults show a complete tail throughout July
while average 3cy michahellis drop the central tail-feathers by the end
of July.
Moult in the wing
coverts:
Normally, adult michahellis do not
include wing-coverts in the partial pre-breeding moult in spring. Consequently, the
coverts start to show extensive wear by June; and by early July feathers
can be very abraded. The feathers most vulnerable for wear are the upper
tertials, the inner coverts and the rear lower scapulars. As can be seen
in some of the images, e.g. image 12 and 10, some tertials and greater
coverts have the top half strongly reduced, leaving only a protruding
shaft-streak in some feathers.
By July, some adults still have present all the
abraded coverts, showing no gaps in the covert rows (see
e.g. image 5). Other
adults start moult, dropping the inner greater coverts and innermost
and outermost median coverts. Adults with new median coverts can be
found at Le Portel by early July as can be seen in image 4.
The lesser covert moult sequence is more
difficult to fully understand as feathers of this tract are dropped
randomly. The start of greater covert moult is pretty obvious, as suddenly
the white-tipped secondaries can be seen from underneath the greater
coverts. Normally, those secondaries are hidden completely.
By the end of
July, the vast majority of adult michahellis have started
wing-covert moult and may look very messy: the old retained feathers are
very abraded and quite some feathers have been dropped, resulting in
various moult gaps.
Timing and
strategy of body & covert moult in July
In line with other large white-headed gulls
in NW Europe, most obvious body feather moult starts with the head
feathers around the eye and the flank feathers (see image 5). From the end of June and
throughout July, feathers are dropped in the area around the eye and
leaves the skin naked. The red orbital ring becomes even more obvious.
Most other body feathers are retained and especially the flank feathers
may look very abraded.
See the Topography
Section for explanation of feather tracts.
Tables:
Adult michahellis remaining
old primaries, July 11 2002 Etaples, NW France. n = 66, m = 6.3, SD
= 1.03 |
P4 /
<P5 old |
4 |
P5
old |
7 |
P6
old |
29 |
P7
old |
19 |
P8
old |
6 |
P9
old |
1 |
P4
/ <P5 old: at least P5-P10 are still old and these scores are
added by the definite P4-P10 birds. Those <P5 birds refer to
sleeping adults. |
adult michahellis remaining
old and fully grown new primaries, July 28 2001 Etaples. |
|
p? |
p3
new |
p4
new |
p5
new |
p6
new |
n
= |
p6
old |
1 |
|
|
|
|
1 |
p7
old |
14 |
2 |
4 |
|
|
20 |
p8
old |
14 |
3 |
5 |
4* |
|
26 |
p9
old |
|
|
|
1 |
1 |
2 |
p10
old |
|
|
|
|
1 |
1 |
m
(average): 2.4 remaining old primaries, SD = 0.69
p?: many birds sleeping, fully grown primaries hidden by tertials, so
often p4 or less. The survey was done at resting and preening adults; presumed sub-adults
(with at least black central markings on the outer primary coverts) were
left out of the score.
*: including two birds with p5 longer than p4, but still not fully
grown (also called p4+). |
adult michahellis remaining
old and fully grown new primaries, July 29 2001 Etaples. |
|
? |
p2
new |
p3
new |
p4
new |
p5
new |
n
= |
p7
old |
16 |
1 |
2 |
3* |
|
22 |
p8
old |
20 |
|
|
8** |
3*** |
31 |
p9
old |
2 |
|
|
|
1 |
3 |
m
(average) = 2.3 remaining old primaries; SD (standard deviation
value) = 0.58.
?: many birds sleeping, fully grown primaries hidden by tertials, so
often p4 or less.
*: including one bird with p4 longer than p3, but still not fully
grown (also called p3+).
**: one bird with p4 longer than p3 but still not fully grown (also
called p3+).
***: one bird with p5 longer than p4 but still not fully grown (also
called p4+). |
adult michahellis scores: Lignano, Venice July 27 - 29 2001. |
prim*
score |
18 |
19 |
20 |
21 |
22 |
23 |
24 |
25 |
26 |
27 |
28 |
29 |
30 |
31 |
32 |
3 |
5 |
4 |
9 |
8 |
19 |
33 |
28 |
46 |
22 |
13 |
5 |
8 |
9 |
4 |
|
|
n:
216, m: 25.3, SD: 2.88 |
fully**
grown |
p0 |
p3 |
p4 |
p5 |
p6 |
|
|
|
|
|
|
|
|
|
|
6 |
9 |
75 |
114 |
18 |
|
|
|
|
|
n:
222 |
m:
4.53 |
SD:
1.02 |
|
|
About
75% is missing the complete median covert bar. |
About
25% is missing / moulting tail-feathers. |
*: prim score =
primary moult score, see Topography for explanation. However, in
this table a different score was used, with shed feathers scoring
0. So, both old feathers and recently shed feathers score 0, fully
grown feathers score 5, still the primary score runs from 0 to 50.
**: Note: 6 bird did not miss a single primary. Those are left out
the primary score. |
This
table presents the results of a research done in the same period at
Lignano, near Venice. As can be read from the table, adults in this region are ahead in primary
moult-score
compared to the birds from NW France. In line with what can be expected,
moult in the rectrices is advanced as well, with about 25% moulting the
tail-feathers. The moult of rectrices starts about a week or 10 days later
at Le Portel (in the first week of August).
|
Yellow-legged
Gull (michahellis) adult, July 09 2010, Maagan Michael, Israel. Picture: Gal Shon. Complete moult. |
Yellow-legged
Gull (michahellis) adult, July 09 2010, Maagan Michael, Israel. Picture: Gal Shon. Complete moult. |
michahellis adult July 28 2010, Malaga, Spain.
Picture: G. Martin. |
Yellow-legged
Gull (michahellis) adult, July 09 2011, Tel Aviv - University Garden, Israel. Picture: Amir Ben Dov.
Mixed pair of michahellis x armenicus. |
michahellis adult. |
photo
1: michahellis adult, July 11 2002, Etaples, NW France. From
unknown origin, ringed green, blue, metal on left leg. |
photo
2: michahellis adult, July 11 2002, Etaples, NW France. The
typical adult which is slightly behind in primary moult: P5 is still old
and P2 is replaced. This individual is in active tail-feather moult. |
photo
3: michahellis adult, July 11 2002, Etaples, NW France. P6
is still old and P3 is growing. |
photo
4: michahellis adult, July 11 2002, Etaples, NW France.
Typical adult. |
photo
5: michahellis adult, July 11 2002, Etaples, NW France.
All wing-coverts abraded. |
photo
6: michahellis group, July 27 2001, Etaples (France). Wings raising adult
with P7-P10 still old and many outer greater and median coverts shed. |
photo
7: michahellis group, July 27 2001, Etaples (France). An adult with P7-P10 still old, the greater coverts shed and the median coverts recently
replaced. |
photo
8: michahellis group, July 27 2001, Etaples (France). Three sub-adults
left and a wing raising adult with P7-P10 old. |
photo
9: michahellis group, July 27 2001, Etaples (France). Note the difference
in moult progress. |
photo
10: michahellis 5cy?, July 27 2001, Le Portel (France). Note the black
central markings on the primary coverts, a sign of immaturity. |
photo
11: michahellis group, July 27 2001, Etaples (France). Left to right: Two wing
raising adults and one 3cy with P8-P10 old. |
Photo
12: michahellis adult, July 7 2001, IJmuiden, the Netherlands. |
photo
13: michahellis adult, July 30 2001, Le Portel (France). P8-P10 still old
and p6 just over P5. |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|