Herring Gull- Zilvermeeuw (argentatus & argenteus)

(last update: December 15, 2011 )

Herring Gull adults February

Introduction

This website deals with the Herring Gulls of the taxa argentatus and argenteus. Argenteus is an abundant breeding species on Iceland, Britain, NW France, Belgium, the Netherlands and N Germany. The taxon argentatus breeds in NE Europe, from Denmark, the coast of the Baltic Sea, Scandinavia and coast of the White Sea. In the centre of the breeding range (Denmark and the NW German coast), birds show intermediate characteristics, indicating there is a cline in argenteus-argentatus. Argenteus in Western Europe is pale-backed, slightly smaller and with relatively much black in the wing-tip; argentatus in the NE Europe is larger, slightly darker on upper-parts and show less black in the wing-tip.
Due to the large breeding range, there is much intra-specific variation in argentatus, with most large, powerful birds in the northern populations, showing the most darkest upper-parts, longest wing and most white in the outer-wing. In this region, some populations may hold substantial number of yellow-legged argentatus. (referred to as "Finnmark yellow-legged argentatus"). In adult plumage, the black sub-terminal spot on P5 is often lacking or strongly reduced to a diffuse streak, broken at the centre. Sometimes P6 lacks black as well. 
In the Baltic region, other yellow-legged argentatus populations occur ("Baltic yellow-legged argentatus"), with adults showing even brighter yellow legs in summer and the upper-parts only slightly darker than West European argenteus, still much paler than Yellow-legged Gull (michahellis). Those Baltic argentatus often show a black sub-terminal spot on P5 in adult plumage.
On these web-pages, we largely follow P.J. Grant: "Gulls, a guide to identification" and E.K Barth's publication as Contribution No. 86, Zoological Museum, University of Oslo: "The circumpolar systematics of Larus argentatus and Larus fuscus with special reference to the Norwegian populations".

adult Herring Gulls - primary pattern

The differences between typical western argenteus from the U.K. and northern Scandinavian argentatus is rather straightforward on primary pattern P5-P10. However, there is a gradient and overlap in features of birds from populations in northern Germany, Denmark and southern Norway. 
Identification is even more challenging in birds from Baltic populations, which show both characteristics of argenteus and argentatus, but sometimes also of the southern taxa michahellis and particularly cachinnans.
Adult birds from the western populations argenteus show more black in the outer primaries, compared to northern argentatus. In argenteus, P5 normally shows small black sub-terminal markings, concentrated on the outer-web as a clear-cut black angular spot and in some birds extending on the inner-web as a diffuse streak. There may be some black on P4 as well, although this might be a clue for near-adult birds. In argentatus, especially from far northern Scandinavia, the black marking on P5 is completely lacking or is very limited and diffuse.
Argenteus
often has a black sub-terminal band on the top of P10, dividing the tip from the mirror. If the black sub-terminal band on P10 is broken, there is often still black on the inner and outer-web's edge of P10. Normally argentatus lacks sub-terminal markings on the outer-web of P10.

Baltic argentatus often have the black marking on P5 confined to the outer-web as a single rectangular small black square, with well-defined borders (not diffuse as in most Finnmark birds). In most Baltic birds, the white tip of P10 fully merges with the mirror, without a single sign of a sub-terminal band. The greyish-white 'tongue' on the inner-web is often broad and continues far down P10, almost creating a thayeri pattern on P10. The shape of the division line between greyish tongue and black outer-web can be an identification clue: In cachinnans this division line often makes a strong angular curve and has a smooth straight line, unlike the pattern of Herring Gulls, which often show a "saw-blade" motive or otherwise much irregularities along the division line. Sometimes wear of the outer primaries may be useful as a first indication of origin: An early moulting species as cachinnans on average show more abrasion in the feathers than later moulting argentatus.

Extensive research on the P10 pattern of Scandinavian argentatus was done for Norwegian birds by the Norwegian ornithologist Edvard Barth, and is presented in the table below. 

Table 4 from E.K. Barth, The circumpolar systematics of Larus argentatus and Larus fuscus with special reference to the Norwegian populations, 1968. 200 Norwegian birds. (Notation converted to what we use in the texts, MM).
locality sample size tip pattern P10 tip pattern P9 colour of P10 inner-web black present from P10 to:
1 2 3 4 1 2 3 4 5 1 2 3 5 6 P7 P6 P5 P4
East Finnmark 15 n 10 1 2 2 5 4 1 1 4 5 7 3 12 3 - 10 5 -
% 67 7 13 13 33 27 7 7 27 33 47 20 80 20 - 67 33 -
Tamsöy in Central Finnmark 33 n 17 7 6 3 10 10 6 0 7 12 20 1 23 10 - 19 13 1
% 52 21 18 9 30 30 18 - 21 36 61 3 70 30 - 58 39 -
Troms 29 n 17 8 3 1 11 9 4 1 4 3 20 6 24 5 - 24 5 -
% 59 28 10 3 38 31 14 3 14 10 69 21 83 17 - 83 17 -
Bodö and Röst 32 n 15 10 3 2 19 3 5 0 3 3 24 5 27 5 1 18 13 -
% 50 33 10 7 63 10 17 - 10 9 75 16 84 16 3 56 41 -
Tarva 20 n 8 7 3 2 11 5 4 0 0 10 8 2 12 8 1 7 10 2
% 40 35 15 10 55 25 20 - - 50 40 10 60 40 5 35 50 10
Stavanger and Lindesnes 42 n 18 13 4 6 23 12 3 2 2 15 23 4 37 5 - 13 29 -
% 44 32 10 14 55 28 7 5 5 36 55 9 88 12 - 31 69 -
Oslo Fiord 29 n 19 6 3 1 20 1 4 1 2 12 14 3 24 5 - 17 12 -
% 66 21 10 3 71 4 14 4 7 42 48 10 83 17 - 59 41 -
.
1: all white
2: trace of black near tip
3: incomplete black bar near tip
4: complete black bar or black tip
1: broad white band over both webs
2: white spot on both webs
3: white spot on inner web only
4: no white spot present
5: 'thayeri-pattern'
1: like mantle colour
2: slightly lighter than mantle colour
3: markedly lighter than mantle colour
5: relatively sharp border between black and grey
6: gradual transition from black to grey

Especially birds from north Scandinavian Finnmark may show the characteristic 'thayeri' pattern in the outer primaries P9 and P10 (column 5 of tip pattern P9 in the table). This pattern is named after the Thayer's Gull, which often shows a particular pattern in the outer primaries: the mirror and the white tip merge and are connected to the pale inner-web (see this image). In other words: the black medial band of the outer-web doesn't fully reach the edge of the inner-web and there is no sub-terminal band. In Thayer's Gull this pattern may be very obvious and, together with the 'string of pearls' may be indicative for identification. 

Primary moult timing and strategy

By October the majority of adult Lesser Black-backed Gulls moulting the outer primaries. There is a clear difference between presumed birds lingering around and birds which will probably migrate further south quickly. The difference can be found in the primary moult strategy. 

Some adult have a large moulting gap in the central primaries (for example P8-P10 old and only P4 fully grown and P5 still slightly shorter than P4) by the first days of September. It is hard to believe these birds will travel large distances non-stop as such moult gaps are highly inefficient for this migration pattern. Probably these birds, which can be found moulting in a period roughly from late August to late November stay in the area throughout their primary moult. They migrate further south either forced by cold winter or when the primaries moult arrives at the far outer primaries.

The other group is represented by birds with a very small moult gap in the primaries or with a very low primary moult score, indicating that primary moult will probably suspended until arrival at the wintering grounds. 

Tables & Graph

remaining SPRING

 

 

 

 

 

 

 

starting date:

 30 Apr

 07 May

14 May

21 May

04 Jun

11 Jun

30 Jul

week no:

18

19

20

21

23

24

31

0

0

0

0

0

 

 

0

1

0

0

0

0

 

 

0

2

0

0

0

0

 

 

1

3

0

0

0

0

 

 

1

4

0

0

0

0

 

 

37

5

0

0

0

0

 

 

22

6

0

0

0

0

 

 

3

7

3

2

1

2

 

 

0

8

16

22

26

18

1

1

0

9

30

13

16

10

1

 

0

10

66

33

37

10

 

 

0

 

 

 

 

 

 

 

 

n=

115

70

80

40

2

1

64

average REM P:

9.4

9.1

9.1

8.7

8.5

8.0

4.4

95% CID =

0.15

0.23

0.20

0.29

6.35

 

0.17

inverse =

1.98

1.99

1.99

2.02

12.71

 

2.00

Variantie (s2) =

0.68

0.90

0.84

0.83

0.50

 

0.46

SD (s) =

0.82

0.95

0.91

0.91

0.71

 

0.68

SE (sx) =

0.08

0.11

0.10

0.14

0.50

 

0.09

 

 

 

 

 

 

 

 

% class:

 

 

 

 

 

 

 

0

0

0

0

0

0

0

0

1

0

0

0

0

0

0

0

2

0

0

0

0

0

0

2

3

0

0

0

0

0

0

2

4

0

0

0

0

0

0

58

5

0

0

0

0

0

0

34

6

0

0

0

0

0

0

5

7

3

3

1

5

0

0

0

8

14

31

33

45

50

100

0

9

26

19

20

25

50

0

0

10

57

47

46

25

0

0

0

 

 

 

 

 

 

 

 

% control:

100

100

100

100

100

100

100

  Table above, data for the Netherlands (2000-2010), results for spring & summer (aggregated week): number of remaining old primaries in adults.
0 = 0 old primaries left (P10 dropped); 10 = all 10 primaries still old and present.

Above: Scatter Plot for PMS in adult Herring Gulls. Number of remaining old primaries throughout the year. Sample size is 1.329 birds.

new: SPRING

 

 

 

 

starting date:

 30 Apr

07 May 

14 May 

21 May 

week no:

18

19

20

21

0

104

60

66

30

1

7

9

12

7

2

4

1

2

3

3

0

0

0

0

4

0

0

0

0

5

0

0

0

0

6

0

0

0

0

7

0

0

0

0

8

0

0

0

0

9

0

0

0

0

10

0

0

0

0

 

 

 

 

 

n=

115

70

80

40

average NEW P:

0.1

0.2

0.2

0.3

95% CID =

0.08

0.10

0.10

0.20

inverse =

1.98

1.99

1.99

2.02

Variantie (s2) =

0.18

0.16

0.21

0.38

SD (s) =

0.43

0.40

0.46

0.62

SE (sx) =

0.04

0.05

0.05

0.10

 

 

 

 

 

% class:

 

 

 

 

0

90

86

83

75

1

6

13

15

18

2

3

1

3

8

3

0

0

0

0

4

0

0

0

0

5

0

0

0

0

6

0

0

0

0

7

0

0

0

0

8

0

0

0

0

9

0

0

0

0

10

0

0

0

0

 

 

 

 

 

% control:

100

100

100

100

  Table above, data for the Netherlands (2000-2010), results for spring & summer (aggregated week): number of new, fully grown primaries in adults.
0 = no new primaries visible. So either P1 still old, or P1 dropped, but the very tip still not visible.
1 = P1 longest new primary. Range between "at least the tip of P1 is visible" until "tip of P2 still shorter than P1".
2 = P2 longest new primary. 3 = etc.

Above: Scatter Plot for PMS in adult Herring Gulls throughout the year. Longest new primary is classified "fully grown". Sample size is 3.271 birds. Birds scored on NEW primaries.

The first date in our analysis on adult Herring Gulls is 02 May 2006 (week 18 - starting 30 April), on which we already found birds with missing primaries (some birds had already shed 3 primaries). This group of actively moulting birds, 43% of the sample, must have started the complete moult somewhere around mid-April and probably several birds may have started earlier.

The last scoring day has been 03 December, when still 20% of the sampled birds had to grow P10. The final stage of the complete moult for all birds has to be somewhere after mid-December. On 03 December we scored 59 birds with P10 fully grown, 13 birds with P9 fully grown and 2 birds with P8 fully grown. They all had no old primaries left.

The first time a new primary (P1) was visible, was also on 02 May 2006 (week 18 - starting 30 April). In this week, 90% of the sample did not show new primaries, but 9% did. Four birds in the sample of 115 birds alreday had P2 as longest primary (fully grown). As soon as the tip of P1 was visible, this was automatically scored "fully grown", and this was the case in 7 birds. For the other primaries to be scored fully grown, they had to exceed the previous primary in length (so P2 fully grown, only from when it is longer than P1).

LINEAR REGRESSION

Onset of primary moult can best be illustrated by a sinusoid for the first few weeks and last weeks, but there appears to be linear dependency over much of the central moult period.

LINEAR REGRESSION FOR OLD REMAINING PRIMARIES

Complete sample: Linear regression (n = 1329, R2= 0.935, sign: P<0,000)
PMS = -0,379* WEEK + 16,324.

LINEAR REGRESSION FOR NEW FULLY GROWN PRIMARIES

Complete sample: Linear regression (n = 3271 R2= 0.927, sign: P<0,000)
PMS = 0,322* WEEK - 6,071.

 

rings argenteus: U.K.
Herring GullHerring Gull (argenteus) L2PT adult, February 10 2016, Katwijk, the Netherlands.
Herring GullHerring Gull GN14174 12cy (argenteus), February 26 2011, Boulogne/Mer, France. Picture: J-M Sauvage.
1cy argenteus in November, ringed in the UK.Herring Gull (argenteus) A3MM 1CY-6CY, November 2006 - February 2011, Oostende, Belgium.
rings argenteus: Belgium
adult argenteus in February, ringed in Belgium. (60451 bytes)Herring Gull BZAK adult (argenteus), February 18 2004, IJmuiden, the Netherlands (52.27N 04.33E). 
Herring GullHerring Gull (argenteus) CKAF 5CY (4th cycle), February 05 2012, Nieuwpoort, Belgium. Picture: Peter Adriaens.
2cy argentatus in February, ringed in Norway. (75189 bytes)Herring Gull (argenteus) CRAC 4CY-5CY (3rd - 5th cycle), May 2011 - December 2012, Koksijde, Belgium. Picture: Peter Adriaens.
1cy argenteus in November, ringed in Belgium. (25637 bytes)Herring Gull (argenteus) FBAD 1CY,2CY,4CY, 6CY, October 2006 - February 2011, Oostende, Belgium. 
adult argentatus in November, ringed in Estonia. (42082 bytes)Herring Gull (argenteus) FMAW 7CY (6th cycle), February 12 2012, Koksijde, Belgium. Picture: Peter Adriaens. 
adult argentatus in November, ringed in Estonia. (42082 bytes)Herring Gull (argenteus) FPAD adult, February 07 2010, Oostduinkerke, Belgium. Picture: Peter Adriaens. 
adult argentatus in November, ringed in Estonia. (42082 bytes)Herring Gull (argenteus) GCAM adult, January 2010 & January-February 2012, Koksijde - De Panne, Belgium. Picture: Peter Adriaens. 
adult argentatus in November, ringed in Estonia. (42082 bytes)Herring Gull (argenteus) GFAB adult, February 13 2011, Oostende, Belgium. Picture: Peter Adriaens. 
adult argentatus in November, ringed in Estonia. (42082 bytes)Herring Gull (argenteus) KNAT adult, May 2010 & January - February 2012, Belgium. Picture: Peter Adriaens. 
adult argentatus in November, ringed in Estonia. (42082 bytes)Herring Gull (argenteus) KPAM adult, February 2011 & November 2012, Oostende, Belgium. Picture: Peter Adriaens. 
Herring GullHerring Gull H-128166 adult (argenteus), February 27 2010, Boulogne/Mer, France (50.42N,1.34E). Picture: J-M Sauvage.
1cy argenteus in November, ringed in Belgium. (25637 bytes)Herring Gull H-127426 adult (argenteus), February 23 2007, Nieuwpoort, Belgium. Picture Ies Meulmeester.
Herring GullHerring Gull (argenteus) BLB H-98296 adult, February 17 2014, Barneveld dump, the Netherlands. Picture: Theo Muusse.
adult argenteus in February, ringed in Belgium. (70167 bytes)Herring Gull H-68803 adult (argenteus), February 07 2003, Boulogne/Mer, France (50.43N 01.37E).
rings argenteus: the Netherlands
1cy argenteus in November, ringed in Belgium. (25637 bytes)Herring Gull FAJS adult female, February 2014 & October 2015, Katwijk & Westkapelle, the Netherlands.
Herring GullHerring Gull (argenteus) FASC adult, February 17 2014, Barneveld dump, the Netherlands. Picture: Theo Muusse.
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argenteus) FASD adult, December 2015 - February 2016 & January 2018, Noordwijkerhout-Katwijk, the Netherlands.
Herring GullHerring Gull (argenteus) M.AMH adult, February & May 2014, the Netherlands. Picture: Theo Muusse & Maarten van Kleinwee.
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argenteus) K.ANC 8CY-9CY, October 2018 & February 2019, Noordwijk & Katwijk, the Netherlands.
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argenteus) Y.ALB 1CY-8CY, 2009-2016, IJmuiden, the Netherlands. 
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argenteus) Y.BAJ 10CY, February & November 2016, IJmuiden & Katwijk, the Netherlands.
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argenteus) YBAX adult, April 2015 - March 2016, IJmuiden, the Netherlands. 
1cy argenteus in November, ringed in Belgium. (25637 bytes)Herring Gull (argenteus) Y.BDS 1CY-5CY, September 2012 - April 2016, IJmuiden, the Netherlands. Pictures: Maarten van Kleinwee & Mars Muusse.
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argenteus) YBHA adult, February 19 2016 & May 29 2017, IJmuiden & Heemskerk, the Netherlands. 
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argenteus) Y.CNJ adult, February 11 2019, Katwijk, the Netherlands.
2cy argentatus in Jan-Febr, ringed in Russia.Herring Gull (argenteus) D||R adult, October 2018 & February 2019, Scheveningen, the Netherlands.

adult argenteus in October, ringed in the Netherlands. (79042 bytes)Herring Gull 5W 6cy (argenteus), February 09 2012, Boulogne/Mer, France (50.42N,1.34E). Picture: J-M Sauvage.

adult argentatus in November, ringed in Estonia. (42082 bytes)Herring Gull (argenteus) PN adult, February 14 2010, De Panne, Belgium. Picture: Peter Adriaens.

Herring GullHerring Gull P8 (argenteus), February-March 2008, Dordrecht, the Netherlands.

2cy argentatus in February, ringed in Norway. (75189 bytes)Herring Gull (argenteus) 4.4 2CY & 5CY, October - December 2015 & February 2018, IJmuiden, Katwijk & Scheveningen, the Netherlands.
Herring GullHerring Gull (argenteus) 4.U 6CY, February 27 2019, Katwijk, the Netherlands.
Herring GullHerring Gull (argenteus) RU adult, February 17 2014, Barneveld dump, the Netherlands. Picture: Theo Muusse.
adult argenteus in February, ringed in the Netherlands. Herring Gull J= 15cy (argenteus), February 04 2001, IJmuiden, the Netherlands (52.27N 04.33E). Is now ringed YAJP.
Herring GullHerring Gull DA 24-27cy (argenteus, 2009-2010, Leiden, Netherlands.
adult argenteus in October, ringed in the Netherlands. (79042 bytes)Herring Gull E261 5-7cy (argenteus), 2009-2011, Leiden, Netherlands. Pictures: M v Kleinwee.
adult argenteus in February, ringed in the Netherlands. (40468 bytes)Herring Gull 5.139.994 10cy (argenteus), February 04 2002, IJmuiden, the Netherlands (52.27N 04.33E).
adult argenteus in February, ringed in the Netherlands. (78145 bytes)Herring Gull 5.265.873 20cy (argenteus), February 16 2010, IJmuiden, the Netherlands (52.27N 04.33E).
adult argenteus in February, ringed in the Netherlands. (78145 bytes)Herring Gull 5.277.846 16cy (argenteus), February 12 2007, IJmuiden, the Netherlands (52.27N 04.33E).
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull adult (argenteus) NLA 5.343.149 January 2011 & February 2015, IJmuiden, the Netherlands.
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull 5.352.764 9cy (argenteus), February 16 2010, IJmuiden, the Netherlands.
adult argenteus in February, ringed in the Netherlands. (78145 bytes)Herring Gull 5.360.719 6+cy (argenteus), February 04 2002, IJmuiden, the Netherlands (52.27N 04.33E).
Herring GullHerring Gull 5.460.074 4+cy (argenteus), February 19 2011, Zoetermeer, Netherlands.
adult argenteus in February, ringed in the Netherlands. (59154 bytes)Herring Gull 5.xx1.121 adult (argenteus), February 04 2002, IJmuiden, the Netherlands (52.27N 04.33E).
Herring GullHerring Gull argenteus 6.125.871 5CY, 8CY & 9CY, 2012 & 2015-2016, IJmuiden, the Netherlands.
Herring GullHerring Gull argenteus NLA 6.168.143 8CY-9CY, February 2018 & March 2019, Katwijk, the Netherlands.
rings argentatus: Germany
adult argentatus in February, ringed in Germany. (67675 bytes)Herring Gull X994 adult (argentatus), February 07 2003, Boulogne/Mer, France (50.43N 01.37E).
adult argenteus in October, ringed in the Netherlands. (79042 bytes)Herring Gull H277 5cy (argentatus), February 25 2011, Leiden, Netherlands. Pictures: M v Kleinwee.
rings argentatus: Norway
Herring Gull (argentatus) J6971 1CY & 6CY, October 2013 & February 2018, Castricum & Katwijk, the Netherlands.
Herring Gull (argentatus) JN744 adult, February 20 2013, Sandøya, Norway. Picture: Ingar Støyle Bringsvor.
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argentatus) JU670 adult, February 11 2019, Katwijk, the Netherlands.
Herring Gull (argentatus) adult NOS FA20131 February 06 2016, Boulogne-sur-Mer, NW France. Picture: Jean-Michel Sauvage.
rings argentatus: Denmark
adult argentatus in February, ringed in Germany. (67675 bytes)Herring Gull argentatus VALS adult, February 27 2011, Enghaveparken - Copenhavn, Denmark. Picture: Kjeld Tommy Pedersen.
Yellow-legged Herring Gull, trapped and ringed in winter in Denmark.
adult argentatus in February, ringed in Germany. (67675 bytes)Herring Gull argentatus DKC 4298575 3CY & 6CY, March & October 2015; February 2018, Leiden, Katwijk & Scheveningen, the Netherlands.
rings argentatus: Russia
Herring GullHerring Gull (argentatus) KUF7 adult, February 25 2016, Terschelling, the Netherlands (53.24N 5.17E). Picture: Jacob Jan de Vries.
Herring GullHerring Gull KN21 8cy (argentatus), February 05 2009, Barneveld (52.07N, 5.34E), the Netherlands. Picture: Herman Bouman.
Herring GullHerring Gull (argentatus) 1CY-8CY KV48 January 11 & February 11 2013, Boulogne, NW France (picture J.M. Sauvage).
1cy argenteus in August, ringed in Belgium. (90559 bytes)Herring Gull (argentatus) RUM C-730262 6CY-7CY, January - February 2018 & March 2019, Katwijk, the Netherlands. 
Herring GullHerring Gull (argentatus) RUM ES-16625 adult, February 17 2014, Barneveld dump, the Netherlands. Picture: Theo Muusse.
rings argentatus: Lithuania
Herring GullHerring Gull argentatus 2L5C adult, February 03 2014, Kaunas, Lithuania. Picture: Boris Belchev.
unringed adult Herring Gulls
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 14 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 28 2012, Noordwijk, the Netherlands. Note thayeri pattern.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 21 2014, Katwijk, the Netherlands. Note thayeri pattern.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 12 2014, Katwijk, the Netherlands. Note thayeri pattern.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 03 2014, Katwijk, the Netherlands. Note thayeri pattern.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 28 2012, Noordwijk, the Netherlands. Note thayeri pattern.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 25 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 25 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 21 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull (argenteus) adult, February 21 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 02 2016, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 21 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argenteus), February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 14 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 14 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 25 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 27 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 14 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 14 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 25 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 03 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 03 2014, Katwijk, the Netherlands. With pale spots in the black on underside of P10.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 21 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 25 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 03 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 03 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 03 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult (argentatus), February 14 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 03 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull (argentatus) adult, February 03 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 25 2012, Noordwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 03 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands. Bird approaching characteristic set for smithsonianus regarding wing-tip.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 19 2015, IJmuiden, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 19 2015, IJmuiden, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 19 2015, IJmuiden, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 21 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 03 2014, Katwijk, the Netherlands.
3cy argentatus in January. (80207 bytes)Herring Gull adult, February 07 2013, Katwijk, the Netherlands.