glaucoides & kumlieni
(last update: October 12, 2011) |
Coordinators:
Dave Brown (Canada)
Bruce Mactavish (Canada)
Chris Gibbins (Scotland)
Peter Adriaens (Belgium)
Mars Muusse (Netherlands)
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Iceland & Kumlien's Gull sub-adult January - April
Below you'll find a copy of the 1992 article "Recognizable Forms -
Subspecies of the Iceland Gull" by
Ron Pittaway published
in: ONTARIO BIRDS APRIL 1992. The PDF version can be downloaded here: PDF PITTAWAY.
"I" in the text below refers to the original author. If any errors occur in this text, please let me know and mail to marsmuusseatgmaildotcom.
Recognizable Forms -
Subspecies of the Iceland Gull
by
Ron Pittaway published
in: ONTARIO BIRDS APRIL 1992
Introduction
The taxonomy of the Iceland Gull
(Larus glaucoides) is complex and
much debated by ornithologists and
birders. In this account, I hope to
clear up some of the confusion about
the status and identification of
the three Iceland Gull subspecies in
Ontario - the Kumlien's Gull (L.g.
kumlieni, Thayer's Gull (L.g. thayerii), and the nominate "Greenland" race (L.g. glaucoides). Here I follow
Godfrey (19861 and James (19911 who
consider the Thayer's Gull to be a
subspecies [race] of the Iceland Gull.
I refer the reader to Plate 36 and the
excellent discussion on the Iceland
Gull in the revised edition of The
Birds of Canada (Godfrey 1986). (Dr.
Earl Godfrey is an internationally
recognized taxonomist and leading
authority on the Iceland Gull.)
Kumlien's Gull:
Kumlien's Gull, pronounced "KOOM-leans" (Terres 1982). breeds
in the eastern Canadian Arctic and
winters mainly in the Atlantic
provinces and Gulf of St. Lawrence,
with small numbers on the Great
Lakes (Godfrey 1986). Contrary to
what many Ontario birders believe,
Kumlien's is the most common race
of the Iceland Gull in the province.
The impression that it is rarer than
the nominate "Greenland" race (L.g.
glaucoides) has persisted for a long
time. This misconception likely resulted from the fact that the
variable grey markings on the
wingtips of many adult Kumlien's are
difficult to see and can be easily overlooked on distant birds. As well,
immature birds are more frequent in
southern Ontario than adults, and
there has been an assumption that
these "white-winged" immatures are
of the nominate race.
Adult Kurnlien's Gulls have
darker grey mantles than adults of
the nominate race, and adult
Glaucous Gulls (L. hyperboreus), In
fact, many Ontario adult Kumlien's
have mantles almost as dark as
Herring Gulls (L. argentatus) (Richard
Poulin, pers. comrn.). Typical adults,
which have light to moderate
pigmentation in the wingtips, can be
reliably identified to race. However,
beware of the fact that the Nelson's
Gull (Herring X Glaucous hybrid) is
rare, but regular, in Ontario. Some
are similar to Kumlieri's, but are
usually distinguishable by their larger
size and heavier bills.
First winter Kumlien's average
darker than nominate birds. Most are
probably indistinguishable in the field (Godfrey 1986). However, those
somewhat darker than normal
individuals having considerable dark
smudging in the wingtips and darker
tails are probably safely called Kumlieri's, but see the reference to
intergrades under Thayer's Gull. For further discussion and excellent
photographs of Kumlien's Gull, the
reader is referred to the recent article
in Birding by Zimmer (1991).
Thayer's Gull
Thayer's Gull was once
considered a subspecies of the
Herring Gull, and more recently a separate species by the American
Ornithologists' Union (1973). based
primarily on the studies of Smith
(1966). Gaston and Decker (1985),
Godfrey (1986), and Snell (1989) have
reported interbreeding between
Kumlien's and Thayer's Gulls,
contrary to Smith (1966). It is now apparent that the Thayer's Gull
represents the dark extreme of the
Iceland Gull complex (Godfrey 1986).
Thayer's Gulls breed in the
western and high Arctic, and winter
mainly on the West coast. Small
numbers occur in migration and
winter in southern Ontario. Typical
adult and first year Thayer's Gulls
can be distinguished from Kumlien's
Gulls "with considerable confidence
in the field" (Godfrey 1986). Intermediates between Thayer's and
Kurnlien's may represent either
extremes in variation or intergrades ([Gaston and Decker 1985, Godfrey
1986, Zimmer 1991).
Beware of two identification
pitfalls. First, an occasional aberrant
Herring Gull can have a Thayer's-like
wing pattern. The Herring's bright
yellow eyes and yellowish (instead of
reddish) fleshy orbital rings should
serve to distinguish it. Second, some
Herring X Glaucous Gull hybrids
(Nelson's Gull) closely resemble
Thayer's Gulls. Their larger size,
paler mantles, and heavier bills
should distinguish the hybrids. First
year hybrids usually have bicoloured, Glaucous-like bills. Detailed
discussion of Thayer's Gull
identification may be found in
Godfrey (1986), Gosselin and David
(1975), Lehman (1980), and Zimmer
(1990).
"Greenland" Iceland Gull
Godfrey (1986) states that the
nominate race (L.g. glaucoides) is "a
non-breeding visitor, mostly in
winter, to southeastern Canada from
the breeding grounds in southern
Greenland". It is by far the rarest of
the three subspecies in Ontario. Earl
Godfrey (pers. comm.) has confirmed
that there is an adult specimen of the
nominate race from Ontario in the
National Museum in Ottawa. This
specimen was collected on 28
November 1974 at Ottawa by Richard
Poulin. Another small and very pale
first winter bird, collected 5
December 1974 at Ottawa by the
author and Richard Poulin, is "almost
certainly" of the nominate race (Earl
Godfrey, pers. cornm.).
The field identification of
nominate birds in Ontario should be
attempted with extreme caution.
Nevertheless, a classic individual
should be recognizable. For a winter
adult (compared with Kumlien's), the
combination of immaculate primary
tips, smaller size, slighter bill, very
pale Glaucous-like mantle, clear
yellow eyes, and pale spotting
confined to the head and nape add up with reasonable certainty to L.g.
glaucoides. Bruce DiLabio [pers.
cornm.) observed such a classic
nominate adult in direct comparison
with adult Kumlien's and Glaucous
Gulls at the Cornwall Dam on 19
January 1991. First year birds cannot
be identified subspecifically in the field (Godfrey 1986). However, small and very pale first winter birds,
having pale at the base of the bill, are
suggestive of the nominate race. See
Figures 2 and 7 in Zimmer (1991). For additional discussion of
identification, see Godfrey (1986) and
Grant (1986).
Summary
Kumlien's Gull is the most
frequently occurring subspecies of
the Iceland Gull in Ontario. The majority of adults can be reliably
identified to race in the field. Most
first winter birds are probably
indistinguishable from nominate
birds. Thayer's Gull is of regular
occurrence, but somewhat less frequent than Kumlien's Gull.
Typical adult and first winter
Thayer's can be distinguished from
Kumlien's with considerable
confidence in the field. The
occurrence of nominate Iceland Gulls
in the province is supported by a
specimen in the National Museum. However, this subspecies is
extremely rare here, and should be
identified with great caution.
Classic
adult nominate individuals are
recognizable in the field with a high
degree of certainty. Most first winter
birds are not separable in the field
from Kumliens. As a final note on
gull identification, don't be afraid to
say "I just don't know what it is!"
Acknowledgements
1 wish to thank the following who
provided me with much valuable
advice and information: Bill Crins,
Bruce DiLabio, Earl Godfrey, Chris
Lemieux, Bruce Mactavish, Richard
Poulin, and Ron Tozer.
Literature cited
American Ornithologists' Union 1973.
Thirty-second supplement to the American
Ornithologists' Union Check-list of North
American Birds. Auk 90: 411-419.
Gaston, A.J. and R. Decker 1985. Interbreeding
of Thayer's Gull, Larus thayeri, and Kurnlien's
Gull, Larus glaucoides kumlieni, on
Southampton Island, Northwest Territories.
Canadian Field-Naturalist 99: 257-259.
Godfrey, W.E. 1986. The Birds of Canada.
Revised Edition. National Museum of
Canada, Ottawa.
Grant, P.J. 1986. Gulls: A Guide to
Identification. Second Edition. Buteo Books,
Vermillion, South Dakota.
James, R.D. 1991. Annotated Checklist of the
Birds of Ontario. Second Edition. Royal
Ontario Museum, Toronto.
Lehman, P, 1980. The identification of Thayer's
Gull in the field. Birding 12: 198-210.
Smith, N.G. 1966. Evolution of some arctic
gulls (Larus): an experimental study of
isolating mechanisms. Ornithological
Monographs No. 4. American Ornithologists'
Union.
Snell, R.R. 1989. Status of Larus gulls at Home
Bay, Baffin Island. Colonial Waterbirds 12:
12-23.
Terres, J.K. 1982. The Audubon Society
Encyclopedia of North American Birds.
Alfred A. Knopf, New York.
Zimmer, K.J. 1990. The Thayer's Gull complex.
Pp. 114-130. in A Field Guide to Advanced
Birding (K. Kaufman). Houghton Mifflin Co.,
Boston.
Zimmer, K.J. 1991. Plumage variation in "Kumlien's" Iceland Gull. Birding 23:
254-269. |
Iceland Gull (kumlieni) 3rd cycle (4CY) A6 January 24 2013, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Peter Adriaens. |
Iceland Gull (kumlieni) 2nd cycle & 3rd cycle C2 January 2012 & January 2013, St John's, Newfoundland, Canada. Picture: Bruce Mactavish & Peter Adriaens. |
Iceland Gull (kumlieni) E6 2nd, 3rd & 7th cycle (3CY, 4CY, 8CY), February 2013, January 2014 & December 2018, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Lancy Cheng. |
Iceland Gull (kumlieni) H1 1st-2nd-3rd-4th cycle (2CY-4CY), 2013 - 2015, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Lisa de Leon, Lancy Cheng, Alvan Buckley & Peter Adriaens. |
Iceland Gull (kumlieni) H8 1st-2nd-3rd-4th cycle, 2013-2014-2015, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Peter Adriaens, Lancy Cheng & Alvan Buckley. |
Iceland Gull (kumlieni) J4 2nd cycle - 3rd cycle (2CY-4CY), December 2013, December 2014 & January 2015, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Alvan Buckley. |
Iceland Gull (kumlieni) K2 3rd cycle (4CY), January 19 2015, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Alvan Buckley. |
Iceland Gull (kumlieni) K6 2nd-5th cycle (3CY-5CY), March 2015 - February 2016 & December 2017, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Lisa de Leon, Lancy Cheng & Frank King. |
Iceland Gull (kumlieni) M2 1st, 3rd & 4th cycle (2CY & 4CY), 2015 & 2017, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Chris van Rijswijk, Lancy Cheng & Lisa de Leon. |
Iceland Gull (kumlieni) M6 2nd-4th cycle (2CY-5CY), December 2015 - January 2018, St. John's, Newfoundland. Picture: Lisa de Leon & Lancy Cheng. |
pale end birds in Europe, classic glaucoides |
Iceland Gull (glaucoides) 4cy, February 05 2012, Texel, the Netherlands. Pictures: René Pop & Jos van de Berg. |
Iceland Gull (glaucoides) 4cy, January 14 - February 20 2012, Hoek van Holland, the Netherlands. Pictures: Peter Soer, Herman Bouman & Theo Muusse. |
Iceland Gull (glaucoides) sub-adult, February 05 2012, Peterhead, Scotland. Pictures: Chris Gibbins. Iris: class 1 (0%>1% speckling). |
Iceland Gull (glaucoides) 4cy, January 28 2012, Katwijk, the Netherlands. Pictures: Rene van Rossum. |
Iceland Gull (glaucoides) 4cy, January 28 2012, Petten, the Netherlands. Pictures: Koos Bakker. |
Iceland Gull (glaucoides) 4cy, January 07 2012, Petten, the Netherlands. Pictures: Ruud Brouwer. |
Iceland Gull (glaucoides) 4cy, January 22 2012, Fraserburgh, Scotland. Picture: Chris Gibbins. |
Iceland Gull (glaucoides) 4cy, February 05 2012, Peterhead, Scotland. Picture: Chris Gibbins. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4CY, February 09 2012, Peterhead, Scotland. Picture: Carl Baggott. |
Iceland Gull (glaucoides) 4CY, February 09 2012, Peterhead, Scotland. Picture: Carl Baggott. |
Iceland Gull (glaucoides) sub-adult, February 09 2012, Peterhead, Scotland. Picture: Carl Baggott. |
Iceland Gull (glaucoides) 4cy / 5cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland Gull (glaucoides) 4cy/5cy, February 23 2012, Kruiningen, the Netherlands. Pictures: Pim Wolf. |
Iceland Gull (glaucoides) 4cy, January 21 - February 19 2012, Brouwersdam, the Netherlands. Pictures: Pim Wolf, Theo Muusse & Luuk Punt. |
Iceland Gull (glaucoides) 3rd cycle (4CY), January 04 2012, Klaksvík, Faroes. Picture: Silas Olofson. |
Iceland Gull (glaucoides) 4cy, January 22 2012, Drachten, the Netherlands. Pictures: Rudy Offereins. |
Iceland Gull (glaucoides) 4cy, February 08 2012, Peterhead, Scotland. Pictures: Chris Gibbins. |
darker birds, still probably glaucoides |
Iceland Gull (glaucoides) 4cy / 5cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. Limited dark on primaries in immature bird. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson, Dark bird overall, outer-webs not contrastingly darker than greater primary coverts. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. dark patches on outer-webs, no sub-terminal markings. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. Dark bird overall, outer-webs not contrastingly darker than greater primary coverts. |
Iceland Gull (glaucoides) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Iceland / Kumlien's Gull 4cy/5cy, January 23 2012, Neeltje Jans, the Netherlands. Pictures: Pim Wolf. Promising bird: darkest areas on outer primaries. |
Iceland / Kumlien's Gull 4cy, January-February 2012, Peterhead, Scotland. Pictures: Chris Gibbins. |
Iceland / Kumlien's Gull 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. Dark bird overall, outer-webs not contrastingly darker than greater primary coverts. Contrasting tail pattern better suits kumlieni. |
Iceland / Kumlien's Gull 4cy, February 08 2012, Peterhead, Scotland. Pictures: Chris Gibbins. |
|
pale birds, but primary patterns with sub-terminal bands advocate kumlieni |
Kumlien's Gull (kumlieni) 4cy, January 27 2012, Zeebrugge, Belgium. Pictures: Filip de Ruwe. |
Kumlien's Gull (kumlieni) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. |
Kumlien's Gull (kumlieni) 4cy, January 24 2012, St John's, Newfoundland, Canada. Picture: Bruce Mactavish.
|
Kumlien's Gull (kumlieni) 4cy, January 24 2012, St John's, Newfoundland, Canada. Picture: Bruce Mactavish. |
Kumlien's Gull (kumlieni) 4cy, February 12 2012, St John's, Newfoundland, Canada. Picture: Dave Brown. |
Iceland Gull (kumlieni) sub-adult, January 23 2013, Quidi Vidi Lake, St. John's, Newfoundland. Picture: Jan Baert. |
dark outer primaries classic for kumlieni |
Kumlien's Gull (kumlieni) 4cy, January 24 2012, St John's, Newfoundland, Canada. Picture: Bruce Mactavish. |
Kumlien's Gull (kumlieni) 4cy, January 24 2012, St John's, Newfoundland, Canada. Picture: Bruce Mactavish. |
Kumlien's Gull (kumlieni) 4cy, January 24 2012, St John's, Newfoundland, Canada. Picture: Bruce Mactavish. |
Kumlien's Gull (kumlieni) 4cy, February 22 2015, Chicago, IL. Picture: Amar Ayyash. |
Kumlien's Gull (kumlieni) 4cy, 28 February - 04 March 2010, near Reykjavik, SW Iceland. Pictures: Hans Larsson. Dark bird overall, outer-webs of outer primaries contrastingly darker than on inners. |
Kumlien's Gull (kumlieni) 3rd cycle (4CY), January 04 2012, Klaksvík, Faroes. Picture: Silas Olofson. |
probable Glaucous x Kumlien's Gull 1st-3rd cycle (2CY), February 2013 - February 2015, Long Beach, Stratford, CT. Picture: Keith Mueller. Bird that became known as "Mr Longbeach", spending subsequent winters at this location. A bulky powerful bird, in many ways intermediate between Glaucous and Kumlien's, in vocals, in size in direct comparison to Herring Gulls, in flight, characteristics, behaviour, and in 'Gestalt'. |