Great Black-backed Gull (Larus marinus)
(last update: July 12, 2015) |
Coordinators:
Jean-Michel Sauvage (France)
Morten Helberg (Norway)
Nils Helge Lorentzen (Norway)
Mars Muusse (Netherlands)
gbbg rings
gbbg PDF's
gbbg 1cy August
gbbg 1cy September
gbbg 1cy October
gbbg 1cy November
gbbg 1cy December
gbbg 2cy January
gbbg 2cy February
gbbg 2cy March
gbbg 2cy April
gbbg 2cy May
gbbg 2cy June
gbbg 2cy July
gbbg 2cy August
gbbg 2cy September
gbbg 2cy October
gbbg 2cy November
gbbg 2cy December
gbbg 3cy January
gbbg 3cy February
gbbg 3cy March
gbbg 3cy April
gbbg 3cy May
gbbg 3cy June
gbbg 3cy July
gbbg 3cy August
gbbg 3cy September
gbbg 3cy October
gbbg 3cy November
gbbg 3cy December
gbbg sub-ad January
gbbg sub-ad February
gbbg sub-ad March
gbbg sub-ad April
gbbg sub-ad May
gbbg sub-ad June
gbbg sub-ad July
gbbg sub-ad August
gbbg sub-ad September
gbbg sub-ad October
gbbg sub-ad November
gbbg sub-ad December
gbbg ad January
gbbg ad February
gbbg ad March
gbbg ad April
gbbg ad May
gbbg ad June
gbbg ad July
gbbg ad August
gbbg ad September
gbbg ad October
gbbg ad November
gbbg ad December
|
Great Black-backed Gull adult September
By September, adult GBBG are still in active moult, most birds growing the central primaries and dropping. As can be seen in so many large white-headed gulls, approximately when P6 is growing, birds start moulting secondaries and rectrices. Of course the tertials (which in a way could be interpreted as inner "secondaries" as well) are included in the moult process much earlier in the season, but when P6 is growing, a new moult wave starts in the outermost secondary (S1), the feather next to P1. The first phase of the secondary moult progresses slowly, one feather at the time, but when the new outer secondaries are visible, a much larger moult gap soon appears in the central secondaries. At the same time, the moult wave that started in the upper tertials continues as well, and the two moult waves meet approximately at S16.
Rectrices (tail feathers) will be moulted in September as well, this process is centrifugal, starting ate the inner feathers and progressing outwards. The exact location of the last remaining old feather is difficult to interpret, but this is not necessarily R6 (the outermost feather). Leaving R5 as last old feather is a common startegy.
An indication of the primary moult of GBBG, and compared to YLG (michahellis) and HG (argentatus) is presented in the tables below:
Tables:
marinus adult PMS Boulogne-sur-Mer (roof) old vs new primaries.
September 21 2002, Boulogne-sur-Mer, NW France.
|
old vs new |
?? |
p5 new |
p6 new |
p7 new |
P4-P10 old |
1 |
0 |
0 |
0 |
P7-P10 old |
2 |
0 |
0 |
0 |
P8-P10 old |
3 |
16 |
0 |
0 |
P9-P10 old |
0 |
15 |
3 |
0 |
P10 old |
0 |
4 |
37 |
0 |
0 old |
0 |
0 |
0 |
1 |
n = 82
m old = 1,8 SD old = 1,08
m new = approx 5,4 SD new =
approx 0,65 |
.
marinus age scored against fully grown new primaries, September 21 2002, Boulogne-sur-Mer, NW France (on the roof).
|
. |
p5 new |
p6 new |
p7 new |
p8 new |
p9 new |
p10 new |
m = |
n = |
SD = |
2nd cy |
- |
- |
2 |
6 |
2 |
- |
8,0 |
10 |
0,67 |
3rd cy |
- |
- |
7 |
1 |
- |
- |
7,1 |
8 |
0,35 |
4th cy |
- |
1 |
3 |
- |
- |
- |
6,8 |
4 |
0,25 |
adult |
35 |
40 |
- |
- |
- |
- |
5,4 |
82 |
0,65 |
.
adult michahellis scores of fully grown new primaries: Etaples, NW France, September 21 2002. Sample and control sample. |
new prims |
I - Sept 21 2002 |
II - Sept 21 2002 |
p7 |
34 |
58 |
p8 |
47 |
50 |
n: |
81 |
108 |
m: |
7.6 |
7.5 |
SD: |
0.50 |
0.50 |
Notes:
All old primaries are shed by late September, except in three birds (2 in sample I, 1 in sample II, all with p7 new);
Due to the very similar plumage of some near-adult michahellis, it's not completely sure all scored birds are indeed fully adult. However, the time lag between these two age-classes is very limited and it's believed only few sub-adults are included at most. |
.
adult michahellis fully grown new primaries, September 29 2001 Etaples |
. |
p5 new |
p6 new |
p7 new |
p8 new |
p9 new |
p10 new |
m = |
n = |
SD = |
morning |
- |
- |
2 |
44 |
2 |
- |
8.0 |
48 |
0.29 |
afternoon |
- |
- |
13 |
95 |
11 |
- |
8.0 |
119 |
0.45 |
.
michahellis age scored against fully grown new primaries, September 29 2001 Le Portel (morning score). |
. |
p5 new |
p6 new |
p7 new |
p8 new |
p9 new |
p10 new |
m = |
n = |
SD = |
2nd cy |
- |
- |
- |
- |
1 |
6 |
9.9 |
7 |
- |
3rd cy |
- |
- |
- |
2 |
7 |
1 |
8.9 |
10 |
0.57 |
4th cy |
- |
- |
- |
3 |
3 |
- |
8.5 |
6 |
0.55 |
adult |
1 |
- |
4 |
20 |
3 |
- |
7.9 |
28 |
0.76 |
.
michahellis age scored against fully grown new primaries, September 29 2001 Le Portel (on the roof). |
|
p7 new |
p8 new |
p9 new |
p10 new |
m = |
n = |
SD = |
2nd cy |
- |
- |
5 |
12 |
9.9 |
17 |
- |
3rd cy |
- |
2 |
16 |
5 |
9.1 |
23 |
0.55 |
4th cy |
4 |
8 |
3 |
- |
7.9 |
15 |
0.70 |
adult |
8 |
26 |
3 |
- |
7.9 |
37 |
0.54 |
|
Great
Black-backed Gull (Larus marinus) J7MZ November 2007 & September 2013 & 2017, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) 6cy J7RH September 10 2011, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) 6cy J9K5 September 24 2007, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) 5cy J9RJ September 02 2010, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) 9cy J90N September 17 2011, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) JE4H / NOS 3005094 2006-2017, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) JE9H 14CY, September 08 2016, Boulogne-sur-Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) JH5R September 2007, August 2010 & September 2012, Boulogne sur Mer, France (50.44 N - 01.35 E).
Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) 5cy JHX9 September 02 2010, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) 6cy JL8C September 01 2010, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) 9cy JP36 September 17 2010, Westkapelle, The Netherlands. Picture Ies Meulmeester. |
Great
Black-backed Gull (Larus marinus) 9cy JZ1K September 17 2011, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) 5cy JZ2J September 24 2007, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) 5cy JZP7 September 24 2004, Scheveningen, the Netherlands.
Primary P6 is fully grown and P9-P10 still old. |
Great Black-backed Gull (L. marinus) J78Z 2CY, 5CY, 11CY, November 2008, November 2011 & September 2017, the Netherlands & France. |
Great
Black-backed Gull JZ11 (Larus marinus) 8+cy, September 10 2011, Boulogne/Mer, France. Picture: J-M Sauvage. |
Great Black-backed Gull (Larus marinus) J031W 7CY, September 11 2017, Boulogne sur Mer, France. Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) 5cy JA112 2008, 2011 & 2012, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull JE165 (Larus marinus) 6+cy: August - September 2012, Boulogne/Mer, France. Picture: J-M Sauvage. |
Great
Black-backed Gull (Larus marinus) 4CY-5CY JE446 2012 & 2013, Boulogne sur Mer, France. Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) 2cy-5cy JH373 2011, 2012 & 2014, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull JH402 (Larus marinus) 4+cy: September 22 2011, Westkapelle, the Netherlands. Picture: M v Kleinwee. |
Great Black-backed Gull (Larus marinus) JH677 8+cy, September 20 2015, IJmuiden, The Netherlands. |
Great Black-backed Gull (Larus marinus) JH844 7CY, August 21 2016, Boulogne sur Mer, France. Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) JP755 6CY, September 11 2017, Boulogne sur Mer, France. Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) adult JP804 September 22 2014, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) JW209 2011-2014, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) JY283 1cy-2cy & 6cy, October 2012 - September 2017, Boulogne sur Mer, France. Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) JY431 4CY-5CY, July 2015 & September 2016, Boulogne sur Mer, France. Picture Jean-Michel Sauvage. |
Great Black-backed Gull (marinus) NOS 354859 25-29CY, August-November 2012-2016, Boulogne-sur-Mer, NW France. Picture: Jean-Michel Sauvage. |
Great
Black-backed Gull (Larus marinus) JHX9 / NOS 3005933 2010, 2012 & 2013,
Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great
Black-backed Gull DKC
3063710 (Larus marinus) adult, September 30 2010, Boulogne/Mer, France. Picture: J-M Sauvage. |
Great
Black-backed Gull NOS
360820 (Larus marinus) 7cy, September 11 2011, Boulogne/Mer, France. Picture: J-M Sauvage. |
Great Black-backed Gull NOS
367624 (Larus marinus) adult: September 18 2004, IJmuiden, the Netherlands (52.27N - 04.33E).
Primary P7 is fully grown while P10 still has to be shed. |
Great Black-backed Gull (Larus marinus) NOS 3020801 8CY, September 22 2017, Boulogne-sur-Mer, France. Picture: Jean-Michel Sauvage. |
Great Black-backed Gull (L. marinus) NOO OA13312 September 24 2006, IJmuiden, the Netherlands. Picture: Ruud Altenburg. |
Great Black-backed Gull (Larus marinus) SVS 9156957 adult, September 30 2016, Westkapelle, the Netherlands. |
Great Black-backed Gull (Larus marinus) SVS 9159239 5cy-9cy, March 2012-January 2016, Boulogne-sur-Mer, France. Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) 3CY-6CY, 8CY DKC 3092175 2012-2017, Boulogne sur Mer, France.
Picture Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) 2E2 adult, April 2012 - September 2015, Sandy Point Beach - Plum Island. Picture: Steve Arena, Jorge de Leon, Billy Clifford & Jeffrey Offermann. |
Great Black-backed Gull (Larus marinus) 5K3 adult, September 29 2010, Gloucester, MA. Picture: Joey Ciaramitaro. |
Great Black-backed Gull (L. marinus) 7F8 7th pre-basic moult (7CY), July - September 2015, Barnstable County, Massachusetts. Picture: Steve Arena. |
Great
Black-backed Gull (Larus marinus) -- adult, September 2011 - October 2017, Boulogne sur Mer, France (50.44 N - 01.35 E).
Picture Jean-Michel Sauvage. |
Great Black-backed Gull (L. marinus) BTO J-0075 8CY-11CY, 2013-2016, Boulogne-sur-Mer, France. Picture: Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus) BTO E-08641 14CY, 18CY-21CY, September-October, Boulogne sur Mer, France (50.44 N - 01.35 E). Picture Jean-Michel Sauvage. |
Great Black-backed Gull (marinus) adult GK5T September 06 2012, Boulogne-sur-Mer, NW France. Picture: Jean-Michel Sauvage. |
Great Black-backed Gull (Larus marinus), adult, September 20 2015, IJmuiden, the Netherlands. |
Great Black-backed Gull (Larus marinus), adult, September 20 2015, IJmuiden, the Netherlands. |
Great Black-backed Gull (Larus marinus), adult, September 20 2015, IJmuiden, the Netherlands. |
Great Black-backed Gull (Larus marinus), adult, September 20 2015, IJmuiden, the Netherlands. |
Great Black-backed Gull (Larus marinus), adult, September 20 2015, IJmuiden, the Netherlands. |
Great Black-backed Gull (Larus marinus), adult, September 24 2015, IJmuiden, the Netherlands. |
Great Black-backed Gull (Larus marinus), adult, September 24 2015, IJmuiden, the Netherlands. |
Great
Black-backed Gull (Larus marinus) adult: September 23 2011,
Scheveningen, the Netherlands. |
Great
Black-backed Gull (Larus marinus) adult: September 23 2011,
Scheveningen, the Netherlands. |
Great
Black-backed Gull (Larus marinus) adult: September 30 2001, Le
Portel - Boulogne sur Mer, France (50.42 N - 01.34
E).
Group of Great Black-backed Gulls and
Herring Gulls (L. argentatus) at Le Portel. |