third calendar year: May
Last update of
this page: 10-10-05 06:32
Introduction:
During
winter and especially early spring, 3cy LBBG have undergone a partial moult
in the winter quarters. This moult typically involved the upper-parts, the
under-parts and the head. Also, the bare parts have developed to a more adult-like
coloration. By May, the coverts and tertials often still consist of old feathers
(retained from previous autumn), which strongly contrast with the renewed
grey upper parts (and sometimes the median covert row). However, not every
3cy LBBG follows this sequence, and we have encountered many returning birds with a variable amount of new tertials,
coverts and even rectrices, and 3cy LBBG with a few new primaries
(arrested moult) as well. It's still preliminary to classify LBBG with
arrested moult in the primaries and allocate them to certain populations.
First figures indicate both intermedius from Scandinavia and local
breeding Dutch intergrades may arrest moult in 3cy in spring.
Detailed moult description:
During
the month of May an unknown number of 3cy birds return to the colony. The
first arrive around April 15th, the majority around May 15th. There may be
good reasons for 3cy birds to return (lack of food supply on the wintering
grounds, to learn the best feeding grounds around the colony, exploit best
breeding places, etc.), but there are risks as well to take on a long
journey back to the breeding grounds; especially if a bird will not start
breeding yet in a particular year. The strongest argument must be the
opportunity to feed for more hours a day compared to areas around the
equator (although LBBG feed at sea at night as well).
3cy graellsii visit the colony in May, but numbers are low. Some
3cy birds stay at NW France, at dumping sites and do not return to the
actually breeding sites, although it's only a short distance.
Most 3cy LBBG stay at the edge of the colony. A theory says the brown
feathers in 3cy birds may protect them against the furious attacks of
territory holding adult males. These brown feathers may be signs for
breeders that they do not participate in mating. Nevertheless, severe fighting
is not uncommon. Still, 3cy birds
daily return to the colony and seem to watch the steps of the elder birds.
They may also predate on eggs, young rabbits, etc in the colony. In May, moult activity is very low.
The
moult of primaries:
Birds
start shedding the innermost primaries. At the end of May all local birds at
least show a gap of P1 and P2, some birds have already shed P4. Adult and
sub-adult LBBG will start their primary moult from mid-June, dropping P1 and
P2.
Compared to 3cy birds, they have a delay of one or two primaries. New
third
generation primaries in 3cy LBBG are adult-like in having grey centers
and white tips, contrasting with the old plain black-brown second generation primaries. Remarkably, from the second half of May 'new' birds
arrive at the colony, showing delayed primary moult compared to average
3cy. Out of ten birds seen on the last day of May, four birds still had
all the primaries present.
Compared
to older birds, the third generation primaries show: smaller tips, P9
lacks a mirror, P10 has a small sized mirror and P3 and P4 show more black
marks (often resulting in a complete sub-terminal band on P4, where adults
only have black on the outer web).
Nearly always, P10 shows obvious white shafts, when seen from below. This
creates a broad white centre in a pale brown feather, on the underside of
P10. Except for the primaries, there are almost no other feathers shed.
The
moult of tail-feathers:
Some
birds start moulting the tail, but May is very early. Most
commonly seen at this age now, is a worn second generation tail,
i.e. a tail with a lot of black, often a broad black tail-band.
Only a few birds are actively moulting some tail-feathers, starting with
the innermost pair. This pair is on top and often easiest bleached and the
tips worn away. Remember that about 30% of the 3cy birds moulted one or
more pairs last winter, resulting in a blocked pattern of white and
non-white tail-feathers. This winter-moult involves tail feathers
randomly, creating an unpredictable black and white tail in spring.
The
moult of coverts:
Most
3cy birds still show brownish second generation wing-coverts (greaters
and lessers at least) while upper-parts and bare parts look very
adult-like. In
May, moult in the coverts is not found in birds staying near the colony.
Birds outside the colony may already start moulting at the end of May. New coverts
are easily recognized since this third generation feathers are
typically uniformly grey, like in adults. Some third generation
feathers however (moulted in the previous autumn like the medians, mantle and
scapulars) may be like second generation feathers in pattern, but those
moulted in summer are much more adult-like. The old remaining second generation feathers always show at least some brown in the centre and show
white fringes, but the exact pattern gets lost by wear.
More about arrested and suspended
moult.
The average moult stage for second summer LBBG
can be seen in the pictures. But some odd individuals turn up as well, for
instance with fresh white tips on P1-P7, while P8-P10 are still second generation (and
P10 obviously
lacks a white mirror). It seems that, in both graellsii and intermedius,
arrested moult is a rare phenomenon, but may occur. See image
30-35.
In his article about identification of fuscus
LBBG
(Birding World, 1998), Lars Jonsson writes about 3cy birds:
"Second-summer fuscus is the most likely to be seen as a
vagrant... Overall, they are much blacker than the corresponding plumages
of graellsii and intermedius and they show much more
adult-looking wing coverts. Also, fuscus very typically shows a
staggered or incomplete renewal of the greater coverts, where a few brown
feathers are retained in an otherwise rather blackish row. On the whole,
the more complete renewal of the wing coverts during the pre-breeding
moult is markedly different to that of intermedius and graellsii,
which typically return for their second summer with a new dark grey
'saddle' but old wing coverts (although quite often the median row will
have been renewed, forming a grey band on the middle of the wing)."
And further: "I would positively identify a second-summer Lesser
Black-backed Gull in May showing predominantly dark wing coverts and very
fresh inner six to eight primaries as a fuscus."
Although this all is very true for fuscus,
it may be an over-simplified analysis of what variation 3cy intermedius
may show when they return in May. Most pictures shown here, probably show graellsii
LBBG or Dutch intergrade LBBG as they were taken near or in
colonies in the Netherlands. But on several occasions, groups of true
migrants from Scandinavia can be found at the beach and several
"puzzling" individuals were encountered of which a few are shown
here. It seems that at least one of the key features to look for (i.e.
arrested moult) occurs in intermedius as well and at least an estimated
4-10%
of the 3cy LBBG age-class may show this features in spring. And this may
be a low estimation, since birds with arrested moult are exactly the birds
"in a hurry" and do not take long rests at the stop-over points. It's
interesting to see in what way fuscus-like features have to
accumulate to make identification as fuscus acceptable. Another web page
is enlarging upon and dedicated to several fuscus-features and occurrence of these
features in graellsii/intermedius.
When intermedius shows
arrested moult in the primaries, combined with advanced moult in the upper-parts, these birds can be mistaken for fuscus. At Le Portel,
France, three such LBBG were present early May 2001, including a very
dark, gentle (female) intermedius. The bird's characteristics
strongly approached those said to be diagnostic for fuscus. Very
dark (blackish) gentle (female?) second summer intermedius, which show
very advanced moult in the upper-parts, combined with arrested moult in
the primaries do occur and approach fuscus in
both coloration and general structure. Now we have come across a few LBBGs,
showing third generation P1-P6 or P1-P7, it's
hard to select diagnostic features to distinguish between the two.
Arrested moult or suspended
moult?
In discussions about arrested moult, suspended moult is
included as well. Strictly, the difference between arrested and suspended
moult lies in the continuation:
Suspended Moult: A bird starts moulting the flight feathers in the
ordinary way (from P1 outwards). At a certain point, the moult stops (is
arrested, e.g. when migration starts). After arrival, the bird catches up
moulting the rest of the flight feathers, which can be recognized as
fresher.
Arrested Moult: A bird starts
moulting the flight feathers in the ordinary way. At a certain point, the
moult stops. After arrival, the bird starts moulting P1 again, thus
doesn't continue moulting the rest of the flight feathers.
In some cases moult continuation is combined and two moult waves start:
one wave at P1 and one wave at the place where moult was arrested, prior
to migration. This is visible as two gaps in the wing (beware of secondary
moult in the outer secondaries (S1, S2, S3).
Arrested moult: in autumn or spring?
It may be important in identification to know whether moult was
arrested prior to migration (in autumn) or moult was arrested later on in
winter, prior to northbound migration. Whatever, it might be very
difficult to judge the correct age of flight feathers by their abrasion and wear. In this discussion, it must be said
that there is a
difference between pigment fading (black feather parts fade to
brown, but grey stays grey) and bleaching (all feathers parts,
including the paler parts turn pale). Where this discussion ends is still
open.
Dutch 3cy LBBG at the Maasvlakte
Do the birds we
see in the Netherlands represent average Dutch LBBG and do they truly
mirror characteristics found in average LBBG? There are about 25.000 breeding
pairs at the Maasvlakte. The average reproduction is 1 young: 25,000 1cy birds leave the colony.
Already 50% die in the first year (12,500 left) and another 25% die in the second year.
Hence, after two years only 9,000 birds remain. If
you were able to count all colonies present on the Maasvlakte
and surroundings in one moment, you would still miss many birds
(1,000's). From
the ringed 300 a year, only 9 where seen again the following year (by only
4 observers). But another year later, around 16 birds were recovered, then
in their 3cy.
Many seem to spend the summer south, where there is no rivalry over nesting
places and enough food supply. Others simply linger around, at sea or at
other feeding stations like dumping sites.
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