Yellow-legged
Gull - Larus michahellis
(last update: October 08 2013) |
Coordinators:
Delfín González
Gabriel Martín
Antonio Gutierrez
Amir Ben Dov
Mars Muusse
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Larus michahellis - adult September
This
website deals with the Yellow-legged Gull taxon michahellis, which
is a common migrant from July to December in NW Europe. After extensive expansion
of the breeding population during the last three decades, it nowadays can
be found breeding in Belgium, the Netherlands and Britain in mixed couples
with both Herring Gull (argenteus) and Lesser Black-backed Gull (graellsii).
There are subtle differences between the populations from the
Mediterranean, Atlantic coast of Portugal and Morocco and from the islands
in the Atlantic. Most pronounced differences can be found in the taxon atlantis,
now regarded as full species by some authors and birds along the Atlantic coast of the Iberian peninsular: lusitanius. Both atlantis and lusitanius are treated in their own sections on this website.
General
description
September is one of the best months to
study michahellis at Le Portel and Etaples, NW France. At the
beach of Le Touquet, near Etaples, large groups congregate and use
the beach as a resting place. They frequent the Dannes landfill as
feeding place and many depend on the local fish-market of Etaples
for food supply. The ports of Boulogne-sur-Mer and the beach of Le
Portel are other good spots to find adult Yellow-legged Gulls.
See Le
Portel page for locations.
The difference between sub-adult
and adult birds is sometimes very difficult. We used the plain grey
greater primary coverts as criteria in our scores. For details on
differences between michahellis and cachinnans, see
e.g. the article by R. Klein & D.
Gruber in Limicola, April 1997.
Moult of remiges and rectrices in
September
All the old primaries have been shed and
new central secondaries grown in almost simultaneously. By the end of the month
the average adult michahellis show eight new fully grown primaries and
an almost completed secondary moult: S1 to ±S8 are fully grown, the
central secondaries are growing and the inner
secondaries are still missing. The tail-feathers
are fully grown or moult is nearly completed: the outer
rectrices which still have to grow an inch or so.
In this section, two tables are included
with the results of primary moult scores from Le Portel. In the first
table, the SD value is strongly influenced by the observation of a single
adult showing P5 as the last fully grown primary. Leaving this single
score out the survey, the SD value would be 0.52, which is more in line
with results from the previous months and the survey from the same day
that is presented in the second table. After leaving the odd single bird
out, the average primary moult score by the end of the month is 7.9 with n
= 64 and SD = 0.53. As can be seen in the table, adults and near-adults
are about one feather behind compared to 3CY birds and about two primaries
behind compared to 2CY birds.
Moult in the wing-coverts
The complete wing-covert moult is in
progress throughout August and is finished by September. The ordinary
moult strategy and sequence started in the outer median coverts, the
innermost medians and innermost greater coverts, the upper tertial and the
outer lower lesser coverts. By mid-August, the median and lower lesser
coverts have been replaced and adult michahellis are missing the
outer greater coverts and some lesser coverts. By late-August, the outer
greater coverts have been replaced as well and the gap in the greater
coverts will be closed at greater coverts #7-#8 in birds following the
ordinary moult strategy, see e.g. here (missing greaters) and here (closing the gap at #7-#8). Moult in the central greater covert is completed by mid September and the
fresh feathers still show a delicate pale grey fringe (see e.g. here).
By early-September, the most advanced birds have replaced all the
wing-coverts and may start again a new additional cycles of wing-covert
moult: the partial wing-covert moult in autumn. They normally start at the
oldest wing-coverts, i.e. the median coverts and lower lesser coverts. See
e.g. here.
The tertial moult is finished by the end of September. Normally the upper
two tertials show wear in the tips while the next four tertials are very
fresh with broad white tips.
The last feathers in the lowest row of
lower scapulars are growing in and by the end of September the complete
scapular and mantle region is pretty neat without moult gaps in the
average adult michahellis, as can be seen in the images from the
end of September (e.g. here, missing one
central lower scapular).
Timing and
strategy of body moult in September
The winter head becomes more obvious in
August and September, where streaking is concentrated around the eye, on the ear coverts
and to the nape, sometimes running down in the neck. Isolated small dark
spots can also be found in the lower neck. Moult of the flank
feathers was started already at the end of July. By August, feathers in
the breast and vent are moulted as well. All under-parts are snowy white
lacking the fluffy loose feathers which were visible in the summer
months.
See the Topography
Section for explanation of feather tracts.
Naked
parts
Adult michahellis shows limited
black on the bill. The colour of legs and bill hardly turns duller than in
full summer plumage. The iris is clear yellow lacking pigmentation; the orbital ring coral
red.
Tables
adult michahellis scores of fully grown new primaries: Etaples, NW
France, September 21 2002. Sample and control sample. |
new
prims |
I
- Sept 21 2002 |
II
- Sept 21 2002 |
p7 |
34 |
58 |
p8 |
47 |
50 |
n: |
81 |
108 |
m: |
7.6 |
7.5 |
SD: |
0.50 |
0.50 |
Notes:
All old primaries are shed
by late September, except in three birds (2 in sample I, 1 in sample
II, all with p7 new);
Due to the very similar plumage of some near-adult michahellis,
it's not completely sure all scored birds are indeed fully adult.
However, the time lag between these two age-classes is very limited
and it's believed only few sub-adults are included at most. |
adult michahellis body
feather moult score, autumn 2001, Maasvlakte, the Netherlands (51.58
N, 4.02 E). |
|
mantle |
us |
ls |
lc |
llc |
mc |
gc |
tt |
R |
31
Aug* 2001 1. |
- |
- |
- |
- |
- |
new |
inners
new |
- |
- |
2. |
- |
- |
- |
- |
- |
new |
many
missing |
- |
- |
3. |
- |
- |
- |
many
missing |
new |
new |
all
visible coverts new |
- |
- |
4. |
- |
- |
- |
many
missing |
- |
new |
many
missing |
- |
- |
5. |
- |
- |
- |
- |
- |
new |
#1-3
new, centrals missing |
- |
- |
03
Sept 2001 1. |
60%
new |
few
centrals new, many missing |
|
new |
#1-3
new, centrals missing |
|
|
Notes:
us = upper scapulars, ls = lower scapulars, lc = lesser coverts, llc
= lower lesser coverts, mc = median coverts, gc = greater coverts,
tt = tertials, R = rectrices.
coverts and tertails are counted from the body to the wingtip.
*: colour-ringed left green EV65, recorded 31 August, 01 September,
03 September 2001. It showed few brown marks on some outer primary
coverts. |
adult michahellis fully grown new primaries, September 29 2001 Etaples |
. |
p5
new |
p6
new |
p7
new |
p8
new |
p9
new |
p10
new |
m
= |
n
= |
SD
= |
morning |
- |
- |
2 |
44 |
2 |
- |
8.0 |
48 |
0.29 |
afternoon |
- |
- |
13 |
95 |
11 |
- |
8.0 |
119 |
0.45 |
michahellis age scored against fully grown new primaries, September 29 2001 Le
Portel |
. |
p5
new |
p6
new |
p7
new |
p8
new |
p9
new |
p10
new |
m
= |
n
= |
SD
= |
2nd cy |
- |
- |
- |
- |
1 |
6 |
9.9 |
7 |
- |
3rd cy |
- |
- |
- |
2 |
7 |
1 |
8.9 |
10 |
0.57 |
4th cy |
- |
- |
- |
3 |
3 |
- |
8.5 |
6 |
0.55 |
adult |
1 |
- |
4 |
20 |
3 |
- |
7.9 |
28 |
0.76 |
michahellis age scored against fully grown new primaries, September 29 2001 Le
Portel (on the roof). |
|
p7
new |
p8
new |
p9
new |
p10
new |
m
= |
n
= |
SD
= |
2nd cy |
- |
- |
5 |
12 |
9.9 |
17 |
- |
3rd cy |
- |
2 |
16 |
5 |
9.1 |
23 |
0.55 |
4th cy |
4 |
8 |
3 |
- |
7.9 |
15 |
0.70 |
adult |
8 |
26 |
3 |
- |
7.9 |
37 |
0.54 |
|
Yellow-legged
Gull michahellis U9AA adult, September 04 2012, Ma'agan Michael, Israel. Picture: Amir Ben Dov.
|
Yellow-legged Gull (michahellis) 19CY-25CY C 53597Boulogne-sur-Mer, NW France. Picture: Jean-Michel Sauvage. |
Yellow-legged
Gull michahellis ITB CA7915 7CY, September 02 2008, Boulogne-sur-Mer, Pas-de-Calais, France. Photo: Jean-Michel Sauvage. |
Yellow-legged Gull michahellis 4CY & 9CY INLD October 2001, September & November 2006, Chioggia, Venezia, Italy.
Picture: Marco Basso & Menotti Passarella. |
Yellow-legged Gull michahellis 8-9CY ITLP 2006-2007, Portalban, Switzerland. Picture: Stephane Aubry. |
Yellow-legged
Gull lusitanius ZDM0 8CY, September 23 2008, Pantín, Spain. Picture: Antonio Gutierrez. |
Yellow-legged Gull lusitanius ZNJM 8CY, September 22 2008, Pantín, Spain. Picture: Antonio Gutierrez. |
Yellow-legged Gull michahellis 9CY HES 875102 September 22 2005, Hauterive, Switzerland. Picture: Stephane Aubry. |
Yellow-legged Gull michahellis 9CY HES 875115 September 01 2005, Hauterive, Switzerland. Picture: Stephane Aubry. |
Yellow-legged Gull michahellis 6CY HES 885446 September 05 2005, Hauterive, Switzerland. Picture: Stephane Aubry. |
Yellow-legged Gull michahellis 5CY HES 885690 September 05 2005, Hauterive, Switzerland. Picture: Stephane Aubry. |
Yellow-legged Gull michahellis 13CY S314 September 11 2011, Poreč landfill, Croatia. Picture: ringing team Luka Jurinovic. |
Yellow-legged Gull michahellis adult S2AU September 24 2010, Lake Balaton, Hungary. Picture: Szabolcs Gál. |
Yellow-legged Gull michahellis DER JC35580 adult, September 02 2010, Veen-Aalburg, the Netherlands. Picture: Bas Verhoeven. |
Yellow-legged Gull michahellis 5CY H-132902 September 02 2009, Westkapelle, the Netherlands. Picture: Ies Muelmeester. |
Yellow-legged Gull michahellis 12CY-13CY M+ August 2013 & September 2014, Neer-Rijkel, the Netherlands. Picture: Jeroen Nagtegaal & Herman Smits. |
Yellow-legged
Gull michahellis J.N adult, May & September 2013, Wijk bij Duurstede & Wageningen, the Netherlands. Picture: PieterGeert Gelderblom & Merijn Loeve. |
Yellow-legged
Gull michahellis DB 5CY-9CY, 2011-2015, the Netherlands. Picture: Maarten van Kleinwee, Merijn Loeve & Mars Muusse. |
Yellow-legged
Gull michahellis adult, September 29 2011, Ashdod, Israel. Picture: Amir Ben Dov. |
Yellow-legged Gull michahellis adult, September 23 2012, Varna, Bulgaria. Picture: Pepa Kostova. |
Yellow-legged Gull michahellis adult, September 24 2011, Porto Ceresio, Lombardy, Italy. Photo: Mirko Tomasi. |
Yellow-legged
Gull michahellis adult, September 02 2010, Eforie Sud, Romania. Picture: Cristian Mihai. |
Yellow-legged Gull michahellis adult, September 20 2011, Ma'agan Michael, Israel. Picture: Amir Ben Dov. |
Yellow-legged Gull michahellis adult, September 20 2011, Ma'agan Michael, Israel. Picture: Amir Ben Dov. |
Yellow-legged
Gull michahellis adult and LBBG, September
21 2002, Dannes, France. |
Yellow-legged
Gull michahellis adult and LBBG, September
21 2002, Dannes, France. |
Yellow-legged Gull michahellis adult, September 21 2006, Portalan, Switzerland. Picture: Stephane Aubry. |
Yellow-legged Gull michahellis adult, September 21 2006, Portalan, Switzerland. Picture: Stephane Aubry. |
Yellow-legged Gull michahellis adult, September 27 2005, Portalban, Switzerland. Picture: Stephane Aubry. |
Yellow-legged Gull michahellis adult, September 29 2006, Portalan, Switzerland. Picture: Stephane Aubry. |
Yellow-legged
Gull michahellis adult, September 22 2002, Etaples, France. |
Yellow-legged
Gull michahellis adult, September 22 2002, Etaples, France. |
Yellow-legged
Gull michahellis adult, September 30 2001, Dannes, France. |
Yellow-legged
Gull michahellis adult, September 30 2001, Dannes, France. |
Yellow-legged
Gull michahellis adult, September 30 2001, Dannes, France. |
Yellow-legged
Gull michahellis x fuscus adult, September 22
2002, Boulogne-sur-Mer, France. |